2002-2005: Post-doctoral fellow at the lab of Kai Simons at Max-Planck Institute for Molecular Cell Biology and Genetics, Dresden, Germany. 2000-2002: Post-doctoral fellow at the lab of Sergio Grinstein at Sickkids Hospital, University of Toronto, Toronto, Canada. 2000: PhD in Pharmacy from University of Coimbra and University Paul Sabatier, Toulouse, France, under the supervision of Leonor Almeida and Robert Salvayre. 1993: Graduated in Pharmaceutical Sciences from the University Coimbra, Portugal