Mar. 2003: PhD in Graduate School of Agriculture, Kyoto University in plant molecular biology for a thesis entitled “Gene identification and characterization of sex chromosome Y in the liverwort, Marchantia polymorpha”, supervised by Prof. Dr. Kanji Ohyama Apr. 2003 to Oct. 2006: Postdoc in Prof. Christopher J. Leaver’s lab in University of Oxford, focusing on plant mitochondria using Arabidopsis thaliana as a model. Nov. 2006: Assistant Professor in Graduate School of Biostudies, Kyoto Univ. started Marchantia researches. Apr. 2013: Associate professor (-2020 Feb.), Professor (from 2020 Mar.) in Graduate School of Science Kobe University. Expanding plant EvoDevo study using Marchantia as a model.