Climate‐induced vegetation change may be delayed in the absence of disturbance catalysts. However, increases in wildfire activity may accelerate these transitions in many areas, including the western boreal region of Canada. To better understand factors influencing decadal‐scale changes in upland boreal forest vegetation, we developed a hybrid modeling approach that constrains projections of climate‐driven vegetation change based on topo‐edaphic conditions coupled with weather‐ and fuel‐based simulations of future wildfires using Burn‐P3, a spatial fire simulation model. We evaluated eighteen scenarios based on all possible combinations of three fuel assumptions (static, fire‐mediated, and climate‐driven), two fire‐regime assumptions (constrained and unconstrained), and three global climate models. We simulated scenarios of fire‐mediated change in forest composition over the next century, concluding that, even under conservative assumptions about future fire regimes, wildfire activity could hasten the conversion of approximately half of Alberta's upland mixedwood and conifer forest to more climatically suited deciduous woodland and grassland by 2100. When fire‐regime parameter inputs (number of fire ignitions and duration of burning) were modified based on future fire weather projections, the simulated area burned was almost enough to facilitate a complete transition to climate‐predicted vegetation types. However, when fire‐regime parameters were held constant at their current values, the rate of increase in fire probability diminished, suggesting a negative feedback by which a short‐term increase in less‐flammable deciduous forest leads to a long‐term reduction in area burned. Our spatially explicit simulations of fire‐mediated vegetation change provide managers with scenarios that can be used to plan for a range of alternative landscape conditions.

2020 is the year of wildfire records. California experienced its three largest fires early in its fire season. The Pantanal, the largest wetland on the planet, burned over 20% of its surface. More than 18 million hectares of forest and bushland burned during the 2019–2020 fire season in Australia, killing 33 people, destroying nearly 2500 homes, and endangering many endemic species. The direct cost of damages is being counted in dozens of billion dollars, but the indirect costs on water‐related ecosystem services and benefits could be equally expensive, with impacts lasting for decades. In Australia, the extreme precipitation (“200 mm day −1 in several location”) that interrupted the catastrophic wildfire season triggered a series of watershed effects from headwaters to areas downstream. The increased runoff and erosion from burned areas disrupted water supplies in several locations. These post‐fire watershed hazards via source water contamination, flash floods, and mudslides can represent substantial, systemic long‐term risks to drinking water production, aquatic life, and socio‐economic activity. Scenarios similar to the recent event in Australia are now predicted to unfold in the Western USA. This is a new reality that societies will have to live with as uncharted fire activity, water crises, and widespread human footprint collide all‐around of the world. Therefore, we advocate for a more proactive approach to wildfire‐watershed risk governance in an effort to advance and protect water security. We also argue that there is no easy solution to reducing this risk and that investments in both green (i.e., natural) and grey (i.e., built) infrastructure will be necessary. Further, we propose strategies to combine modern data analytics with existing tools for use by water and land managers worldwide to leverage several decades worth of data and knowledge on post‐fire hydrology.

The timing, extent, and severity of forest wildfires have increased in many parts of the world in recent decades. These wildfires can have substantial and devastating impacts on water supply, ecohydrological systems, and sociohydrosystems. Existing frameworks to assess the magnitude and spatial extent of these effects generally focus on local processes or services and are not readily transferable to other regions. However, there is a growing need for regional, continental, and global scale indices to assess the potential effect of wildfires on freshwater availability and water supply resilience. Such indices must consider both the individual and compound effects of wildfires. In so doing, this will enable comprehensive insights on the water security paradigm and the value of hydrological services in fire‐affected areas around the globe.