1996 Received Ph.D. in Virology at Graduate School of Medicine (under the supervision of Prof. Kazuyoshi Ikuta), Hokkaido University, Hokkaido, Japan. (also supervised by Prof. Akio Adachi at Kyoto University, 1994-1996) 1996-1998 Research Resident at National Institute of Infectious Diseases, Tokyo, Japan 1998-1999 CNRS Research Fellow at Institute Pasteur (Nobel Prize-awarded Prof. Françoise Barré-Sinoussi’s lab), Paris, France 1999-2001 Duke Postdoctoral Fellow at Duke University Medical Center (Prof. Bryan Cullen’s lab), NC, USA 2001-present Principal Investigator at National Institute of Infectious Diseases, Tokyo, Japan 2002-2003 JICA Expert of Virology at Noguchi Memorial Institute for Medical Research, University of Ghana, Accra, Ghana 2009-2017 Adjunct Associate Professor at Tokyo Medical and Dental University, Tokyo, Japan 2010-2013 Invited Associate Professor at Osaka University, Osaka, Japan 2018-present Visiting Professor at University of Yamanashi, Integrated Graduate School, Yamanashi, Japan 2019-present Visiting Professor at Kumamoto University, Joint Research Center for Human Retrovirus Infection, Kumamoto, Japan