Whereas there is evidence that mixed-species approaches to production forestry in general can provide positive outcomes relative to monocultures, it is less clear to what extent multiple benefits can be derived from specific mixed-species alternatives. To provide such insights requires evaluations of an encompassing suite of ecosystem services, biodiversity, and forest management considerations provided by specific mixtures and monocultures within a region. Here, we conduct such an assessment in Sweden by contrasting even-aged Norway spruce (Picea abies)-dominated stands, with mixed-species stands of spruce and birch (Betula pendula or B. pubescens), or spruce and Scots pine (Pinus sylvestris). By synthesizing the available evidence, we identify positive outcomes from mixtures including increased biodiversity, water quality, esthetic and recreational values, as well as reduced stand vulnerability to pest and pathogen damage. However, some uncertainties and risks were projected to increase, highlighting the importance of conducting comprehensive interdisciplinary evaluations when assessing the pros and cons of mixtures.

The performance of growth models implemented in the Swedish Forest Planning System Heureka was evaluated. Four basal area growth models were evaluated by comparing their predictions to data from five-year growth records for 1711 permanent sample plots of the National Forest Inventory (NFI-data). Also, two alternative implementations of Heureka, including a combined stand- and tree-level basal area growth model and a single tree-level model, respectively, were evaluated using data from 57 blocks in a thinning experiment (GG-data) involving Scots pine ( L.) and Norway spruce ( (L.) Karst) in which the trees were monitored for 30 years after the first thinning. The predicted volume growth was also compared to observed values. Growth models based on data from 1970’s and 1980’s overestimated growth in the NFI test plots from the early 2000’s by about 3%. Stand-level models had larger precision than tree-level models. Basal area growth was underestimated in dense NFI-plots and overestimated in non-thinned GG-plots, illustrating an un-solved modelling problem. Basal area growth was overestimated by 2–5% also in the GG-plots over the whole observation period. Volume growth was however accurately predicted for pine and underestimated by 2% for spruce. The relative prediction error did not increase with increasing length of prediction period. Thinning response models calibrated with GG-data worked well in the total application and produced growth levels for different thinning alternatives in line with observations.Pinus sylvestris Picea abies

Continuous-cover forestry (CCF) has been recognized for the production of multiple ecosystem services, and is seen as an alternative to clear-cut forestry (CF). Despite the increasing interest, it is still not well described how CCF would affect the carbon balance and the resulting climate benefit from the forest in relation to CF. This study compares carbon balances of CF and CCF, applied as two alternative land-use strategies for a heterogeneous Norway spruce (Picea abies) stand. We use a set of models to analyze the long-term effects of different forest management and wood use strategies in Sweden on carbon dioxide emissions and carbon stock changes. The results show that biomass growth and yield is more important than the choice of silvicultural system per se. When comparing CF and CCF assuming similar growth, extraction and product use, only minor differences in long-term climate benefit were found between the two principally different silvicultural systems.

The effects of intensifying the management of 15% of the Swedish forest land on potential future forest production over a 100-year period were investigated in a simulation study. The intensive management treatments, which were introduced over a period of 50 years, were: intensive fertilization of Norway spruce (IntFert); bulking-up Norway spruce elite populations using somatic embryogenesis (SE-seedlings); planting of lodgepole pine, hybrid larch, and Sitka spruce (Contorta, Larch, and Sitka); fertilization with wood ash on peatlands (Wood ash); and conventional fertilization in mature forests (ConFert). Potential sites for applying intensive forest management (IFM) to sites with low nature conservation values were determined with a nature conservation score (NCS). Four different scenarios were simulated: “Base scenario”, which aimed at reducing the negative impact on nature conservation values, “Fast implementation”, “No IntFert” (IntFert was not used), and “Large Forest Companies”, where the majority of plots were selected on company land. Total yields during the 100-year simulation period were about 85–92% higher for the intensive forest management scenarios than for the reference scenario (business as usual). In the “No IntFert” scenario total production was 1.8% lower and in the “Large Forest Companies” scenario total production was 4.8% lower than in the “Base scenario”. “Fast implementation” of IFM increased yield by 15% compared to the “Base scenario”. Norway spruce SE-seedlings and IntFert gave the highest yields, measured as total production during the 100-year simulation period, but relative to the yields in the reference scenario, the highest increases in yield were for Contorta. The “Base scenario” and “No IntFert” gave the highest yields for plots with the lowest NCS, but plots with higher NCS had to be used in the “Fast implementation” and “Large Forest Companies” scenarios. More than half of the effect on future growth of IFM methods was because of increased intensity in the regenerations. It took a relatively long time (40–60 years) for the simulated IFM treatments to result in a significant increase in stem volume production.

In a stand with planted Norway spruce (Picea abies (L.) Karst.) and naturally regenerated birch (Betula pendula Roth and B. pubescens Ehrh.) in southern Sweden, the proportion of birch was reduced in 0.1-ha plots at pre-commercial thinning to 0%, 20% or 50% of the total stem number (2000–2500 stems ha⁻¹ after the thinning). The growth of both species was surveyed 4 and 10 years later. No significant between-treatment differences were observed on either occasion in total yield or in spruce diameter (apart from a more negative skewness of the diameter distribution of Norway spruce in the mixture with 50% birch compared to pure spruce). Birch retained its initial advantage in dominant height (of about 2 m) throughout the 10-year measurement period; hence, height growth rates of Norway spruce and birch were very similar. After three, four and 10 growing seasons, whipping damage to Norway spruce caused by birch was registered in the 20% and 50% birch mixtures, in which the proportions of damaged spruces were found to be similar, with 17% and 15% of damaged trees at the last inventory, respectively. In addition, simulations of the stands’ development from the time of the last measurement until final felling at a total age of 66 years indicated that growth, yield and net present value decrease with increasing birch proportion, but if the proportion of birch removed in subsequent thinnings is increased, the between-treatment differences are reduced.

Four different management strategies were applied to a young mixed stand of Scots pine ( L.), Norway spruce ( (L.) Karst.) and birch ( Roth and Ehrh.) in southern Sweden. All strategies included an initial precommercial thinning to ca. 2400 stems ha. The different aims were: (i) a conifer-dominated stand with focus on high productivity; (ii) a conifer-dominated stand with high quality timber; (iii) keeping a stem-wise species mixture; (iv) a mosaic-wise species mixture. Stem selection according to the different strategies were simulated with a starting point from plots with a 5 m radius. All strategies were applied to all of the plots. A growth simulator was used to simulate the stand development up to final felling. This study illustrates the possibilities for influencing the structure of a mixed stand through precommercial thinning. The study also illustrates the long-term effects on stand structure and volume yield by consequently applying a management strategy from precommercial thinning until final felling. Precommercial thinning was found to be a useful tool to influence the stand structure in accordance to the aims set. However, the opportunities for influencing the stand by precommercial thinning were restricted by natural settings and unconditional considerations (e.g. tree vitality, stem quality, regular spacing), beyond what could be judged from stand average data. The stem volume production during a rotation was 6% lower for (iii) and (iv) compared to (i) and (ii), mainly due to a greater proportion of birch in the former strategies.Pinus sylvestris Pices abies Betula pendula Betula pubescens –1