Based on the experimental estimation of the key dynamic properties of a seven-storey building made entirely of cross-laminated timber (CLT) panels, the finite element (FE) model updating was performed. The dynamic properties were obtained from an input-output full-scale modal testing of the building in operation. The chosen parameters for the FE model updating allowed the consideration of the timber connections in a smeared sense. This approach led to an excellent match between the first six experimental and numerical modes of vibrations, despite spatial aliasing. Moreover, it allowed, together with the sensitivity analysis, to estimate the stiffness (affected by the connections) of the building structural elements. Thus, the study provides an insight into the as-built stiffness and modal properties of tall CLT building. This is valuable because of the currently limited knowledge about the dynamics of tall timber buildings under service loadings, especially because their design is predominantly governed by the wind-generated vibrations.