The geological record contains evidence for numerous pronounced perturbations in the global carbon cycle, some of which are associated with mass extinction. In the Carnian (Late Triassic), evidence from sedimentology and fossil pollen points to a significant change in climate, resulting in biotic turnover, during a time termed the ‘Carnian Pluvial Episode’ (CPE). Evidence from the marine realm suggests a causal relationship between the CPE, a global ‘wet’ period, and the injection of light carbon into the atmosphere. Here we provide the first evidence from a terrestrial stratigraphic succession of at least five significant negative C-isotope excursions (CIE)’s through the CPE recorded in both bulk organic carbon and compound specific plant leaf waxes. Furthermore, construction of a floating astronomical timescale for 1.09 Ma of the Late Triassic, based on the recognition of 405 ka eccentricity cycles in elemental abundance and gamma ray (GR) data, allows for the estimation of a duration for the isotope excursion(s). Source mixing calculations reveal that the observed substantial shift(s) in δ¹³C was most likely caused by a combination of volcanic emissions, subsequent warming and the dissociation of methane clathrates.

The generally arid Late Triassic climate was interrupted by a wet phase during the mid-Carnian termed the Carnian Pluvial Episode (CPE). Quantitative palynological data from the Mercia Mudstone Group in the Wessex Basin (UK) reveal vegetation changes and palaeoclimate trends. Palynostratigraphy and bulk organic carbon isotope data allow correlation to other Carnian successions. The palynostratigraphy indicates that the Dunscombe Mudstone is Julian and the lowest part of the overlying Branscombe Mudstone Formation is Tuvalian. The Aulisporites acme characterizing the CPE in Tethyan successions and the Germanic Basin is missing in the UK. The quantitative palynological record suggests the predominance of xerophyte floral elements with a few horizons of increased hygrophytes. A humidity signal is not seen owing to the dry climate in central Pangea. Also, the signal might be masked by the overrepresentation of xerophyte regional pollen and the predominance of xerophyte hinterland flora. The bias towards regional pollen rain is enhanced by the potential increase in continental runoff related to seasonally humid conditions and differences in pollen production rates and transport mechanisms. The vegetation of British CPE successions suggests a more complex climate history during the Carnian, indicating that the CPE is not recognized by the same changes everywhere. Supplementary material: A detailed lithological log of the Strangman's Cove (Devon) outcrop with the description of the Mercia Mudstone Group lithostratigraphical units, a description of the laboratory techniques, seven photoplates with selected spores and pollen grains, a list of all identified palynomorphs, Excel sheets with the palynological and palynofacies counts, bulk organic carbon isotope ratios and total organic carbon values of the Strangman's Cove outcrop are available at https://doi.org/10.6084/m9.figshare.c.4138085

Palynological and paleobotanical investigation of bonebeds and other strata of the Csehbánya Formation from the vertebrate locality at Iharkút (Bakony Mts, Hungary) reveals well-preserved Santonian palynological assemblages dominated by the Normapolles group, with a minor component consisting of other angiosperm pollen, some gymnosperm pollen, and spores. Eleven species of Normapolles-type pollen grains belonging to seven genera and fruit remains of a new taxon, Sphaeracostata barbackae gen. et sp. nov., are described. The new species is very abundant in the material, represented by ca. 1000 specimens. The genus Caryanthus Friis and an unnamed form previously reported from Haţeg by Lindfors et al. (2010) are also present. Plants producing Normapolles-type pollen grains diversified during the Late Cretaceous, with a bloom in the Santonian. The palynostratigraphy of the Upper Cretaceous terrestrial sediments in the studied region is based on Normapolles-related species. The studied assemblage is assigned to the Oculopollis zaklinskaiae-Tetracolporopollenites (Brecolpites) globosus Zone (or Zone C) indicating a late Santonian age. Comparison of the Iharkút palynoflora with other known Upper Cretaceous palynofloras of Central Europe shows diachronous occurrence of Normapolles taxa at different geographic localities and warrants further investigation. The ecological requirements of the amphibian fauna reflect azonal conditions controlled by the availability of water, which is in agreement with the inferred ecological conditions based on the paleobotanical investigations. The fauna is of entirely non-marine character, further supported by isotope studies, in line with our data showing that the palynological samples contain no marine forms

Fossil plant assemblages including spores and pollen grains provide useful information on past ecosystems and the response of terrestrial biotas to various environmental perturbations. New quantitative palynological data from the Chinle Formation of the American Southwest suggest that a floral turnover occurred in the middle Norian (between 217 and 213 Ma). Analysis of plant communities reveals that this turnover was followed by a complete reorganization of the riparian vegetation, driven by changes in fluvial styles and the tectonic regime of the basin, as well as a gradual transition toward a more arid climate. Marked increases in Klausipollenites gouldii, Patinasporites spp., and Froelichsporites traversei are probable indicators of environmental stress, such as increased aridity, perturbations of atmospheric pCO₂, acid rain, and atmospheric aerosol accumulation due to volcanism in connection with the Pangean rifting and uplift of the Cordilleran arc. Comparison of the vegetation turnover with younger assemblages from the Chinle Formation in New Mexico revealed similar floral turnover patterns, suggesting two distinct drier periods as a result of multiple climatic oscillations. The climate-induced floral turnover may have contributed to the vertebrate faunal turnover as the loss of wetland habitat space and an increase in xerophytic plants may have dwindled the supply of palatable vegetation for herbivores. The onset of the floral turnover in Arizona roughly corresponds to the Manicouagan impact event, but a direct causal link is still speculative.