Rapid climate change and altered disturbance regimes represent increasing stressors to the stability of existing ecosystems. Resilience is a widely used framework for post-disturbance response, but resilient responses are emergent properties resulting from component processes of persistence, recovery, and reorganization, with different mechanisms at work in each mode. We present a model of scaled resilience, which allows resilience to be decomposed across scales of space, time, and levels of biological organization. Using case examples of post-fire resilience in dry conifer forests of interior western North America, we illustrate the increased clarity gained by separating scale-dependent mechanisms of persistence, recovery, and reorganization. We conclude by describing how the scaled resilience framework can be applied in land and fire management by distinguishing relevant management actions before, during, and after wildfire.

Over the past four decades, annual area burned has increased significantly in California and across the western USA. This trend reflects a confluence of intersecting factors that affect wildfire regimes. It is correlated with increasing temperatures and atmospheric vapour pressure deficit. Anthropogenic climate change is the driver behind much of this change, in addition to influencing other climate-related factors, such as compression of the winter wet season. These climatic trends and associated increases in fire activity are projected to continue into the future. Additionally, factors related to the suppression of the Indigenous use of fire, aggressive fire suppression and, in some cases, changes in logging practices or fuel management intensity, collectively have produced large build-ups of vegetative fuels in some ecosystems. Human activities provide the most common ignition source for California’s wildfires. Despite its human toll, fire provides a range of ecological benefits to many California ecosystems. Given the diversity of vegetation types and fire regimes found in the state, addressing California’s wildfire challenges will require multi-faceted and locally targeted responses in terms of fuel management, human-caused ignitions, building regulations and restrictions, integrative urban and ecosystem planning, and collaboration with Tribes to support the reinvigoration of traditional burning regimes.

Climate change is increasing the frequency and extent of high-severity disturbance, with potential to alter vegetation community composition and structure in environments sensitive to tipping points between alternative states. Shrub species display a range of characteristics that promote resistance and resilience to disturbance, and which yield differential post-disturbance outcomes. We investigated differences in shrub patch size and stem density in response to variations in fire severity, vegetation community, and post-disturbance reproductive strategies in Sky Island forested ecosystems in the southwestern United States. Patterns in shrub structure reflect the effects of fire severity as well as differences among species with alternate post-fire reproductive strategies. Increased fire severity correlates with larger patch sizes and greater stem densities; these patterns are observed across multiple fire events, indicating that disturbance legacies can persist for decades. High severity fire produces the largest shrub patches, and variance in shrub patch size increases with severity. High severity fire is likely to promote expansion of shrub species on the landscape, with implications for future community structure. Resprouting species have the greatest variability in patch structure, while seeding species show a strong response to disturbance: resprouting species dominate at low disturbance severities, and obligate seeders dominate high severity areas. Differential post-fire reproductive strategies are likely to generate distinct patterns of vegetation distribution following disturbance, with implications for community composition at various scales. Shrub species demonstrate flexible responses to wildfire disturbance severity that are reflected in shrub patch dynamics at small and intermediate scales.