Long-proboscid scorpionflies from amber reveal the variety and complexity of mid-Cretaceous pollinating insects.

Lara, M.B., Bashkuev, A. & Wang, B., 1.10.2014. Argentinopanorpa miguezi gen. et sp. nov.: first record of Triassic Mecoptera (Permochoristidae) from the Cuyo Basin (Mendoza, Argentina). Alcheringa 39, 000‒000. ISSN 0311 5518 A new genus and species of Permochoristidae, Argentinopanorpa miguezi, is described based on a complete forewing from the Upper Triassic Potrerillos Formation (Mendoza Province, Argentina). The new taxon is placed in the subfamily Agetopanorpinae and distinguished from other genera by the following combination of characters: Rs+MA with six branches, MP with eight branches (MP₁ and MP₃ with additional forks), CuA and MP fused basally for a short distance. This is the first record of Permochoristidae in the Triassic of South America and the first formal description of Mecoptera from the Cuyo Basin. Previously reported taxa attributed to Mecoptera from the Triassic of South America are briefly discussed with notes on their systematic position. María Belén Lara (corresponding author)[lara.maria.belen@live.com.ar], Area Entomología (Facultad de Ciencias Exactas y Naturales y Agrimensura-Universidad Nacional del Nordeste) and Area Paleontología (Centro de Ecología Aplicada del Litoral-Consejo Nacional de Investigaciones Científicas y Técnicas-Universidad Nacional del Nordeste), Casilla de Correo 128, 3400 Corrientes, Argentina; Alexei Bashkuev [fossilmec@gmail.com], Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya ul. 123, Moscow, 117997 Russia; Bo Wang [savantwang@gmail.com], Steinmann Institute, University of Bonn, 53115 Bonn, Germany and State Key Laboratory of Palaeobiology and Stratigraphy; Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China. Received 4.7.2014; revised 19.8.2014; accepted 7.9.2014.

The systematic position of Microptysmatidae and Protomeropidae is critically reviewed in light of the latest reassessment by Minet et al. The Microptysmatidae belongs to a stem-group of Trichoptera, or at least to stem-Amphiesmenoptera (depending on chosen methodology), rather than to its own order Permotrichoptera as suggested by Minet et al., and Protomeropidae are not Mecoptera. The double anal loop of Kamopanorpa Martynov, 1928 is not significantly different from that of more advanced amphiesmenopterans, while in Microptysmella Kukalová-Peck & Willmann, 1990, anal veins show clear tendency to looping (not a “fairly different arrangement”). If Microptysmatidae are considered in the traditional way, the polarity of characters listed by Minet et al. as supporting monophyly of the order Permotrichoptera can be treated just the opposite: divided veins RS1, RS2 and M1 are symplesiomorphies with Protomeropidae, and simple M4 is the synapomorphy of Microptysmatidae, Cladochoristidae, and Amphiesmenoptera s. str. In addition, three new species of Kamopanorpa are described from the Permian deposits of Minusinsk Coal Basin, South Siberia: Kamopanorpa rasnitsyni sp. nov., K. sivchikovi sp. nov., and K. rotunda sp. nov. Some traits of fore- and hindwing venation of Microptysmatidae are discussed.