In complex low-altitude environments, unmanned aerial vehicles (UAVs) require reliable positioning, yet Global Navigation Satellite System (GNSS) signals are vulnerable to occlusion and interference. Pseudolite-augmented cooperative localization, which combines ground base-station signals with inter-UAV relative observations, can complement GNSS in such environments. However, two practical issues remain in real-world deployment: UAV-to-base-station (U-B) and UAV-to-UAV (U-U) observations have markedly different error statistics that a unified noise adjustment cannot handle, and the conservative covariance estimates produced by Covariance Intersection (CI) fusion bias the innovation-based adaptive noise estimation in distributed architectures. To address these issues, this paper proposes a Distributed Group Covariance Compensation Adaptive Kalman Filter (DGCC-AKF) for collaborative enhancement of UAV regional localization. DGCC-AKF establishes a group adaptive mechanism that independently adjusts the noise covariance matrices of U-B and U-U observations, enabling observation-type-level adaptive weighting that suppresses anomalous U-B or U-U measurements at the group level. In addition, a bounded covariance compensation factor is incorporated to alleviate the CI-induced conservatism in the adaptive noise estimation. The proposed method is evaluated on a 2800 km² semi-physical testbed based on the Ground-based High-precision Local Positioning System (GH-LPS) pseudolite network using measured U-B observations and high-dynamic (>300 km/h) flight trajectories collected from a fixed-wing platform across three independent flight sessions. Results demonstrate that under observation fault periods, the proposed method improves 3D positioning accuracy by up to about 75% over single-UAV extended Kalman filter (EKF). Compared with two advanced algorithms in this field, variational Bayesian adaptive Kalman filter (VBAKF) and maximum correntropy criterion Kalman filter (MCC-EKF), it is the only scheme that remains accurate and stable across all UAVs and fault types. The framework provides a practical step toward field deployment for resilient multi-UAV cooperative navigation in pseudolite-augmented GNSS-denied regions.