Herbaceous communities in forest ecosystems on the southern part of the Russian Plain appeared in the Middle Miocene (∼10 Ma BP). In the Late Miocene (∼7 Ma BP), feather-grass steppe associations...

Surface Kastanozem of the Lower Volga area was first studied as a part of the pedocomplex, with the lower part (148–160 cm) formed in Early Khvalynian Chocolate clays (13–15 ka), the middle part (100–148 cm) in a mixed clay-loess sediment sand, and the upper part (0–100 cm) in loess. This resulted from local aeolian transport, with the source material derived from the rewinding of marine sediments. They are enriched in aggregates of Chocolate clays and glauconitic grains of a fine sand-course silt size and have similar contents of clay minerals. The high salinity of similar types evidences marine genesis for both Chocolate clays and source material for loess sediments. Clay fragments of a sand and silt size are responsible for the heavy texture and high gypsum content of loess. The study of soils with the focus on micromorphology and clay mineralogy allows the identification of the complex character of a shift from marine to sub-areal sedimentation. This shift was accompanied by short breaks in sedimentation, allowing the development of synlithogenic soil horizons of Late Khvalynian, after-Khvanynian, and Boreal time. The features of shallowly buried soil horizons confirm increased aridity after the last deglaciation. Surface Calcic Kastanozem is a full Holocene soil reflecting the present environment. However, it is deeply influenced by shallow buried soil horizons and Chocolate clays.

Neanderthals were widespread during the Middle Palaeolithic (MP) across Europe and Asia, including the Caucasus Mountains. Occupying the border between eastern Europe and West Asia, the Caucasus is important region regarding the Neanderthal occupation of Eurasia. On current radiometric estimates, the MP is represented in the Caucasus between about 260–210 ka and about 40 ka. Archaeological record indicates that several culture diverse MP hominin populations inhabited the Caucasus, but the region complex population history during this period remains poorly understood. In this paper, we identify for the first time the archaeological evidence indicating contacts between two culture diverse MP Neanderthal populations in the North Caucasus and discuss the nature of these contacts. Basing on the lithic assemblages that we excavated at Mezmaiskaya cave in the north-western Caucasus (Kuban River basin) and Saradj-Chuko grotto in the north-central Caucasus (Terek River basin), dating from MIS 5 to MIS 3, and comparative data from other MP sites in the Caucasus, we identify two large cultural regions that existed during the late MP in the North Caucasus. The distinctive toolkits and stone knapping technologies indicate that the MP assemblages from Mezmaiskaya cave and other sites in the west of North Caucasus represent a Caucasian variant of the Eastern Micoquian industry that was wide spread in central and eastern Europe, while the assemblages from Saradj-Chuko Grotto and other sites in the east of North Caucasus closely resemble the Zagros Mousterian industry that was wide spread in the Armenian Highlands, Lesser Caucasus and Zagros Mountains. The archaeological evidence implies that two culture diverse populations of Neanderthals settled the North Caucasus during the Late Pleistocene from two various source regions: from the Armenian Highlands and Lesser Caucasus along the Caspian Sea coast, and from Russian plain along the Sea of Azov coast.