Authors’ contribution Study conception and experimental design: I.-C.S. and Y.-K.S.; experiments: V.K.Y., S.A.S., and I.-H.F.; data collation and analysis: I.-C.S., Y.-K.S., H.-Y.C., V.K.Y., S.A.S., and I.-H.F.; manuscript writing: I.-C.S., Y.-K.S., C.-T.Y., H.-C.H., and C.-M.L.; reagents, materials, and experimental infrastructure: H.-Y.C., C.-T.Y., H.-C.H., and C.-M.L. All authors have read and approved the final submitted version of the manuscript. All authors read and approved the final submitted version of the manuscript.