Geographically isolated wetlands (GIWs), those surrounded by uplands, exchange materials, energy, and organisms with other elements in hydrological and habitat networks, contributing to landscape functions, such as flow generation, nutrient and sediment retention, and biodiversity support. GIWs constitute most of the wetlands in many North American landscapes, provide a disproportionately large fraction of wetland edges where many functions are enhanced, and form complexes with other water bodies to create spatial and temporal heterogeneity in the timing, flow paths, and magnitude of network connectivity. These attributes signal a critical role for GIWs in sustaining a portfolio of landscape functions, but legal protections remain weak despite preferential loss from many landscapes. GIWs lack persistent surface water connections, but this condition does not imply the absence of hydrological, biogeochemical, and biological exchanges with nearby and downstream waters. Although hydrological and biogeochemical connectivity is often episodic or slow (e.g., via groundwater), hydrologic continuity and limited evaporative solute enrichment suggest both flow generation and solute and sediment retention. Similarly, whereas biological connectivity usually requires overland dispersal, numerous organisms, including many rare or threatened species, use both GIWs and downstream waters at different times or life stages, suggesting that GIWs are critical elements of landscape habitat mosaics. Indeed, weaker hydrologic connectivity with downstream waters and constrained biological connectivity with other landscape elements are precisely what enhances some GIW functions and enables others. Based on analysis of wetland geography and synthesis of wetland functions, we argue that sustaining landscape functions requires conserving the entire continuum of wetland connectivity, including GIWs.

Efforts are increasingly being made to classify the world’s wetland resources, an important ecosystem and habitat that is diminishing in abundance. There are multiple remote sensing classification methods, including a suite of nonparametric classifiers such as decision-tree (DT), rule-based (RB), and random forest (RF). High-resolution satellite imagery can provide more specificity to the classified end product, and ancillary data layers such as the Normalized Difference Vegetation Index, and hydrogeomorphic layers such as distance-to-a-stream can be coupled to improve overall accuracy (OA) in wetland studies. In this paper, we contrast three nonparametric machine-learning algorithms (DT, RB, and RF) using a large field-based dataset (n = 228) from the Selenga River Delta of Lake Baikal, Russia. We also explore the use of ancillary data layers selected to improve OA, with a goal of providing end users with a recommended classifier to use and the most parsimonious suite of input parameters for classifying wetland-dominated landscapes. Though all classifiers appeared suitable, the RF classification outperformed both the DT and RB methods, achieving OA >81%. Including a texture metric (homogeneity) substantially improved the classification OA. However, including vegetation/soil/water metrics (based on WorldView-2 band combinations), hydrogeomorphic data layers, and elevation data layers to increase the descriptive content of the input parameters surprisingly did not markedly improve the OA. We conclude that, in most cases, RF should be the classifier of choice. The potential exception to this recommendation is under the circumstance where the end user requires narrative rules to best manage his or her resource. Though not useful in this study, continuously increasing satellite imagery resolution and band availability suggests the inclusion of ancillary contextual data layers such as soil metrics or elevation data, the granularity of which may define its utility in subsequent wetland classifications.

Wetlands provide many ecosystem services, including sediment and carbon retention, nutrient transformation, and water quality improvement. Although all wetlands are biogeochemical hotspots, geographically isolated wetlands (GIWs) receive fewer legal protections compared with other types of wetlands because of their apparent isolation from jurisdictional waters. Here, we consider controls on biogeochemical functions that influence water quality, and estimate changes in ecosystem service delivery that would occur if these landscape features were lost following recent US Supreme Court decisions (i.e., Rapanos, SWANCC). We conclude that, despite their lack of persistent surfacewater connectivity or adjacency to jurisdictional waters, GIWs are integral to biogeochemical processing on the landscape and therefore maintaining the integrity of US waters. Given the likelihood that any GIW contributes to downstream water quality, we suggest that the burden of proof could be shifted to assuming that all GIWs are critical for protecting aquatic systems until proven otherwise.