He earned his PhD in Developmental Genomics, with research focused on conserved regulatory regions—such as enhancers—and their roles in heart development and disease, using zebrafish as a model organism. In addition to his work in developmental biology, he has contributed to the fields of ecotoxicology and reproductive health, studying antioxidant responses and stress biomarkers in fish, amphibians, and reptiles. He previously earned an MSc in Biology of Marine Organisms at the European Institute for Marine Studies, where he concentrated on genomic and physiological approaches. His academic journey began with a BSc in Marine Science from the University of Naples Federico II, driven by a strong interest in the biochemical and physiological adaptations of marine organisms.