Education and qualifications: 1954 M.D. University Göttingen, Germany 1956 PhD, University Göttingen, Germany 1963 PhD, Karolinska Institute, Stockholm 1964 M.D. Karolinska Institute, Stockholm 1966 Assistant Professor, Karolinska Institute, Stockholm 1963-1970 Head, Department of Medicine, Swedish Radiation Protection Authority 1970-1993 Professor and Head, Department of Medical Radiobiology, Karolinska Institute and Hospital, Stockholm 1994-2019 Consultant Professor, Department of Oncology and Pathology, Karolinska Institute