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Dr. Marcelo Kovačić
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0 Ecology
0 Ichthyology
0 Taxonomy
0 Gobiidae
0 Mrphology

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Natural History Museum Rijeka Lorenzov prolaz 1 HR-51000 Rijeka Croatia

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Journal article
Published: 22 July 2021 in Journal of Marine Science and Engineering
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Cryptobenthic fishes were often overlooked in the past due to their cryptic lifestyle, so knowledge of their ecology is still incomplete. One of the most poorly studied taxa of fishes in the Mediterranean Sea is clingfish. In this paper we examine the habitat preferences of three clingfish species (Lepadogaster lepadogaster, L. candolii, and Apletodon incognitus) occurring in the Gulf of Trieste (Northern Adriatic). The results show that all three species have a cryptic lifestyle and are well-segregated based on their depth distribution and macro- and microhabitat preferences. L. lepadogaster inhabits shallow waters of the lower mediolittoral and upper infralittoral, where it occurs on rocky bottoms under stones. L. candolii similarly occurs in the rocky infralittoral under stones, but below the lower distribution limit of L. lepadogaster, and in seagrass meadows, where it occupies empty seashells. Such hiding places in seagrass meadows are also occupied by A. incognitus, which mostly occurs below the lower distribution limit of L. candolii. Despite the overlap of depth and macrohabitat, the probability of individuals of two species encountering each other or competing in the same habitat is low when the depth range is combined with the microhabitat preferences of these species.

ACS Style

Domen Trkov; Danijel Ivajnšič; Marcelo Kovačić; Lovrenc Lipej. Factors Influencing Habitat Selection of Three Cryptobenthic Clingfish Species in the Shallow North Adriatic Sea. Journal of Marine Science and Engineering 2021, 9, 789 .

AMA Style

Domen Trkov, Danijel Ivajnšič, Marcelo Kovačić, Lovrenc Lipej. Factors Influencing Habitat Selection of Three Cryptobenthic Clingfish Species in the Shallow North Adriatic Sea. Journal of Marine Science and Engineering. 2021; 9 (8):789.

Chicago/Turabian Style

Domen Trkov; Danijel Ivajnšič; Marcelo Kovačić; Lovrenc Lipej. 2021. "Factors Influencing Habitat Selection of Three Cryptobenthic Clingfish Species in the Shallow North Adriatic Sea." Journal of Marine Science and Engineering 9, no. 8: 789.

Journal article
Published: 05 July 2021 in Zootaxa
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A new cryptobenthic gobiid species Hetereleotris nasoramosa sp. nov. is described based on the holotype and five paratypes collected from the north-eastern part of Socotra Island, Arabian Sea, from moderately large pieces of coral rocks with holes at depths of 8–11 m. Molecular phylogenetic analysis placed the new species within the genus Hetereleotris. Hetereleotris nasoramosa sp. nov., differs from all species of Hetereleotris in having developed tentacles extending from each anterior and posterior nostril and five transverse suborbital papillae rows (instead four or six in other species). The new species superficially resembles the recently described Red Sea endemic species Cerogobius petrophilus by having forward-set, elevated eyes, a short snout, a moderately large mouth, a relatively deep and short caudal peduncle, and developed tentacles on the head, but differs from it by the same characters of developed tentacles extending from each anterior and posterior nostril and five transverse suborbital papillae rows as from other Hetereleotris species. Both species also share a specific habitat preference for tight holes in rock covered by micro-algae. A full description of the species is provided as well as a revised key to the species of Hetereleotris.

ACS Style

Marcelo Kovačić; Sergey V. Bogorodsky; Uwe Zajonz; Luke Tornabene. A new species of Hetereleotris (Teleostei: Gobiidae) from the Socotra Archipelago (north-western Indian Ocean), a rare case of a hole-associated adaptation in gobiid fishes. Zootaxa 2021, 4996, 283 -300.

AMA Style

Marcelo Kovačić, Sergey V. Bogorodsky, Uwe Zajonz, Luke Tornabene. A new species of Hetereleotris (Teleostei: Gobiidae) from the Socotra Archipelago (north-western Indian Ocean), a rare case of a hole-associated adaptation in gobiid fishes. Zootaxa. 2021; 4996 (2):283-300.

Chicago/Turabian Style

Marcelo Kovačić; Sergey V. Bogorodsky; Uwe Zajonz; Luke Tornabene. 2021. "A new species of Hetereleotris (Teleostei: Gobiidae) from the Socotra Archipelago (north-western Indian Ocean), a rare case of a hole-associated adaptation in gobiid fishes." Zootaxa 4996, no. 2: 283-300.

Journal article
Published: 18 June 2021 in Journal of Marine Science and Engineering
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Coralligenous habitat is considered as one of the most important special habitat types in the Mediterranean; however, due to its inaccessibility, little is known about it, although it is considered as one of the Mediterranean’s richest habitats in terms of species. Due to a low number of studies, it was presumed that the richness of coralligenous fish assemblages is underestimated using traditional visual census methods which are not applicable to the deep, steep, and vertical slopes of coralligenous cliffs and do not capture exhaustively cryptobenthic species commonly found in this habitat. This paper aims at producing a more complete assessment of fish assemblages on a coralligenous cliff by combining different methods, particularly the deep vertical transect visual census and square with anesthetics method. A total of 76 fish species were recorded on a single coralligenous cliff, supporting the opinion that coralligenous cliffs are important Mediterranean biodiversity hotspots. The analysis of species traits between species recorded by the different methods showed how complementary they are to better describe species compositions. Hence, the result of this study demonstrates that the combined use of methods is essential for a more exhaustive description of the whole fish community structure and for accurate estimates of the abundance and diversity patterns, particularly in complex habitats such as coralligenous cliffs.

ACS Style

Alen Soldo; Igor Glavičić; Marcelo Kovačić. Combining Methods to Better Estimate Total Fish Richness on Temperate Reefs: The Case of a Mediterranean Coralligenous Cliff. Journal of Marine Science and Engineering 2021, 9, 670 .

AMA Style

Alen Soldo, Igor Glavičić, Marcelo Kovačić. Combining Methods to Better Estimate Total Fish Richness on Temperate Reefs: The Case of a Mediterranean Coralligenous Cliff. Journal of Marine Science and Engineering. 2021; 9 (6):670.

Chicago/Turabian Style

Alen Soldo; Igor Glavičić; Marcelo Kovačić. 2021. "Combining Methods to Better Estimate Total Fish Richness on Temperate Reefs: The Case of a Mediterranean Coralligenous Cliff." Journal of Marine Science and Engineering 9, no. 6: 670.

Journal article
Published: 21 May 2021 in Journal of Marine Science and Engineering
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Data on the distribution and ecology of cryptobenthic fish of marine caves in the Mediterranean Sea are extremely scarce but necessary for scientists and marine managers alike in order to understand these fish’s ecological role and assess their conservation status. Broadscale surveys by implementing underwater visual census and photographic sampling in marine caves of the northeastern Mediterranean Sea, within different expeditions during the last 5 years, brought to light new records of eight rarely reported cryptobenthic fish species. To a smaller extent, complementary citizen science data from diving professionals of Crete were used to fill distribution gaps. A total of 36 new records (66 individuals) from 18 marine caves and caverns of the Aegean and northeastern Levantine Seas were assembled, belonging to the gobies Corcyrogobius liechtensteini, Didogobius splechtnai, Gammogobius steinitzi, and Thorogobius ephippiatus, the blenny Microlipophrys nigriceps, the tripterygiid Tripterygion melanurum, the speleophilic bythitid Grammonus ater, and the gobiesocid Lepadogaster cf. lepadogaster. The above species have been rarely reported from the Eastern Mediterranean Sea, with D. splechtnai and G. steinitzi being recorded for the first and second time from Greek waters, respectively, while L. cf. lepadogaster constitutes the second record of a clingfish species in a marine cave of the Aegean Sea. Interesting behavioral and ecological habits were also noted for some species, based on in situ observations and photographic evidence. Our study contributes to filling gaps in the knowledge of cave fish diversity and demonstrates that cryptobenthic mobile species in understudied cryptic habitats are more common than previously thought in the Mediterranean Sea.

ACS Style

Michail Ragkousis; Markos Digenis; Marcelo Kovačić; Stelios Katsanevakis; Vasilis Gerovasileiou. Rarely Reported Cryptobenthic Fish in Marine Caves of the Eastern Mediterranean Sea. Journal of Marine Science and Engineering 2021, 9, 557 .

AMA Style

Michail Ragkousis, Markos Digenis, Marcelo Kovačić, Stelios Katsanevakis, Vasilis Gerovasileiou. Rarely Reported Cryptobenthic Fish in Marine Caves of the Eastern Mediterranean Sea. Journal of Marine Science and Engineering. 2021; 9 (6):557.

Chicago/Turabian Style

Michail Ragkousis; Markos Digenis; Marcelo Kovačić; Stelios Katsanevakis; Vasilis Gerovasileiou. 2021. "Rarely Reported Cryptobenthic Fish in Marine Caves of the Eastern Mediterranean Sea." Journal of Marine Science and Engineering 9, no. 6: 557.

Journal article
Published: 09 November 2020 in Zootaxa
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A checklist of 73 gobiid species (Teleostei: Gobiidae) recorded to date from the Mediterranean Sea is established following the evidence approach for checklists. The Mediterranean gobiofauna currently has 62 known native species and 11 alien species. An identification key to gobiid species known from the area is provided. The principles of character selection for the key are discussed.

ACS Style

Marcelo Kovačić. Checklist of gobies (Teleostei: Gobiidae) of the Mediterranean Sea and a key for species identification. Zootaxa 2020, 4877, 75 -101.

AMA Style

Marcelo Kovačić. Checklist of gobies (Teleostei: Gobiidae) of the Mediterranean Sea and a key for species identification. Zootaxa. 2020; 4877 (1):75-101.

Chicago/Turabian Style

Marcelo Kovačić. 2020. "Checklist of gobies (Teleostei: Gobiidae) of the Mediterranean Sea and a key for species identification." Zootaxa 4877, no. 1: 75-101.

Journal article
Published: 13 October 2020 in Journal of Marine Science and Engineering
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Gobies (Gobiiformes: Gobiidae) are the most species-rich family of fishes in general, and the most abundant fish group in the European seas. Nonetheless, our knowledge on many aspects of their biology, including the population genetic diversity, is poor. Although barriers to gene flow are less apparent in the marine environment, the ocean is not a continuous habitat, as has been shown by studies on population genetics of various marine biota. For the first time, European marine goby species which cannot be collected by common fishery techniques were studied. The population genetic structure of two epibenthic species, Gobius geniporus and Gobius cruentatus, from seven localities across their distribution ranges was assessed, using one mitochondrial (cytochrome b) and one nuclear gene (first intron of ribosomal protein gene S7). Our results showed that there is a great diversity of haplotypes of mitochondrial gene cytochrome b in both species at all localities. Global fixation indices (FST) indicated a great differentiation of populations in both studied gobies. Our results did not show a geographic subdivision to individual populations. Instead, the data correspond with the model of migration which allow divergence and recurrent migration from the ancestral population. The estimated migration routes coincide with the main currents in the studied area. This matches well the biology of the studied species, with adults exhibiting only short-distance movements and planktonic larval stages.

ACS Style

Katarína Čekovská; Radek Šanda; Kristýna Eliášová; Marcelo Kovačić; Stamatis Zogaris; Anna Maria Pappalardo; Tereza Soukupová; Jasna Vukić. Population Genetic Diversity of Two Marine Gobies (Gobiiformes: Gobiidae) from the North-Eastern Atlantic and the Mediterranean Sea. Journal of Marine Science and Engineering 2020, 8, 792 .

AMA Style

Katarína Čekovská, Radek Šanda, Kristýna Eliášová, Marcelo Kovačić, Stamatis Zogaris, Anna Maria Pappalardo, Tereza Soukupová, Jasna Vukić. Population Genetic Diversity of Two Marine Gobies (Gobiiformes: Gobiidae) from the North-Eastern Atlantic and the Mediterranean Sea. Journal of Marine Science and Engineering. 2020; 8 (10):792.

Chicago/Turabian Style

Katarína Čekovská; Radek Šanda; Kristýna Eliášová; Marcelo Kovačić; Stamatis Zogaris; Anna Maria Pappalardo; Tereza Soukupová; Jasna Vukić. 2020. "Population Genetic Diversity of Two Marine Gobies (Gobiiformes: Gobiidae) from the North-Eastern Atlantic and the Mediterranean Sea." Journal of Marine Science and Engineering 8, no. 10: 792.

Regular papers
Published: 28 September 2020 in Journal of Fish Biology
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The clingfish (Gobiesocidae) genus Gouania (Nardo 1833) is endemic to the Mediterranean Sea and inhabits, unlike any other vertebrate species in Europe, the harsh intertidal environment of gravel beaches. Following up on a previous phylogenetic study, we revise the diversity and taxonomy of this genus, by analysing a comprehensive set of morphological (meristics, morphometrics, micro computed tomography imaging), geographical and genetic (DNA‐barcoding) data. We provide descriptions of three new species, G. adriatica sp. nov., G. orientalis sp. nov., G. hofrichteri sp. nov. as well as re‐descriptions of G. willdenowi (Risso 1810) and G. pigra (Nardo 1827) and assign neotypes for the latter two species. In addition to elucidating the complex taxonomic situation of Gouania, we discuss the potential of this enigmatic clingfish genus for further ecological, evolutionary and biodiversity studies that might unravel even more diversity in this unique Mediterranean fish radiation.

ACS Style

Maximilian Wagner; Marcelo Kovačić; Stephan Koblmüller. Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra. Journal of Fish Biology 2020, 98, 64 -88.

AMA Style

Maximilian Wagner, Marcelo Kovačić, Stephan Koblmüller. Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra. Journal of Fish Biology. 2020; 98 (1):64-88.

Chicago/Turabian Style

Maximilian Wagner; Marcelo Kovačić; Stephan Koblmüller. 2020. "Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra." Journal of Fish Biology 98, no. 1: 64-88.

Journal article
Published: 18 August 2020 in Zootaxa
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Corcyrogobius pulcher sp. nov. is described from off Île de Ngor, Dakar, Senegal. Corcyrogobius pulcher is distinguished from its two congeners by having the rear edge of the jaws ending posteriorly below mideye, second dorsal fin I/9, pectoral fin rays 17, pelvic fins oval or truncated posteriorly, scales in lateral series 26–27, anterior oculoscapular head canal with pore β, suborbital row b of sensory papillae anteriorly beginning below vertical of posterior edge of eye, dark vertical caudal bar, branchiostegal membrane without intense dark spot, cheek with two oblique whitish stripes, the first going from the eye downwards and forward to the posterior jaws, the second on the preopercular, alternating with brown oblique stripe going from behind the eye downwards and forward across the cheek. Furthermore, mitochondrial COI-barcoding data unambiguously support the species-level distinctiveness of the three Corcyrogobius species. A key to the species of Corcyrogobius is provided.

ACS Style

Marcelo Kovačić; Peter Wirtz; Ulrich K. Schliewen. A new species of Corcyrogobius (Teleostei: Gobiidae) from Île de Ngor, Senegal. Zootaxa 2020, 4834, 121 -132.

AMA Style

Marcelo Kovačić, Peter Wirtz, Ulrich K. Schliewen. A new species of Corcyrogobius (Teleostei: Gobiidae) from Île de Ngor, Senegal. Zootaxa. 2020; 4834 (1):121-132.

Chicago/Turabian Style

Marcelo Kovačić; Peter Wirtz; Ulrich K. Schliewen. 2020. "A new species of Corcyrogobius (Teleostei: Gobiidae) from Île de Ngor, Senegal." Zootaxa 4834, no. 1: 121-132.

Journal article
Published: 23 April 2020 in Zootaxa
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The updated checklist of Adriatic Sea fishes with a critical assessment of each species using an evidence approach is provided. Each fish species in Adriatic Sea listed in the last published checklist and those reported in published new records not included in the most recent Adriatic checklist, have been included. Of the total of 466 fish species, the presence in the Adriatic Sea was confirmed for 444 species by at least one positive record of the species in the area, the presence of 10 species is still unconfirmed, and 12 fish species are excluded from the list. An evidence approach protocol is recommended for general use for compiling checklists of marine fishes.

ACS Style

Marcelo Kovačić; Lovrenc Lipej; Jakov Dulčić. Evidence approach to checklists: critical revision of the checklist of the Adriatic Sea fishes. Zootaxa 2020, 4767, 1 -55.

AMA Style

Marcelo Kovačić, Lovrenc Lipej, Jakov Dulčić. Evidence approach to checklists: critical revision of the checklist of the Adriatic Sea fishes. Zootaxa. 2020; 4767 (1):1-55.

Chicago/Turabian Style

Marcelo Kovačić; Lovrenc Lipej; Jakov Dulčić. 2020. "Evidence approach to checklists: critical revision of the checklist of the Adriatic Sea fishes." Zootaxa 4767, no. 1: 1-55.

Journal article
Published: 10 March 2020 in Zootaxa
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Silhouettea ghazalae sp. nov. is described from Qeshm Island, Persian Gulf. Silhouettea ghazalae sp. nov. is distinguished from congeners by having: small mental fold present on chin, head length 31.4-32.4% of standard length, head width 24.5% of standard length, second dorsal fin I/11, anal fin I/13, breast with large cycloid scales, predorsal area naked, suborbital row b anteriorly beginning below anterior edge of pupil, posteriorly ending below pore β, suborbital row c anteriorly extending more than row b and posteriorly extending less than row b, suborbital row cp oblique with four papillae, body with four ill-defined midlateral blotches and the fifth a triangular mark on the caudal fin base, no clearly defined pale saddles on back, and the first dorsal fin pigmented with dots and with dark blotch present anteriorly. A key to Silhouettea species is provided. A dataset including novel and publicly available mtDNA COI sequences of 12 species from the Persian Gulf and Oman Sea gobies belonging to eight genera have been assembled in order to provide a reference dataset for DNA barcoding studies. The new species is further characterised by a minimum K2P distance of 21% to its closest relatives in our dataset, Cabillus tongarevae in the mtDNA COI barcode region.

ACS Style

Marcelo Kovačić; Reza Sadeghi; Hamid Reza Esmaeili. New species of Silhouettea (Teleostei: Gobiidae) from Qeshm Island, Iran and the DNA barcoding of the Persian Gulf and Oman Sea gobies. Zootaxa 2020, 4750, 49 -66.

AMA Style

Marcelo Kovačić, Reza Sadeghi, Hamid Reza Esmaeili. New species of Silhouettea (Teleostei: Gobiidae) from Qeshm Island, Iran and the DNA barcoding of the Persian Gulf and Oman Sea gobies. Zootaxa. 2020; 4750 (1):49-66.

Chicago/Turabian Style

Marcelo Kovačić; Reza Sadeghi; Hamid Reza Esmaeili. 2020. "New species of Silhouettea (Teleostei: Gobiidae) from Qeshm Island, Iran and the DNA barcoding of the Persian Gulf and Oman Sea gobies." Zootaxa 4750, no. 1: 49-66.

Journal article
Published: 03 March 2020 in Scientia Marina
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Cryptobenthic fishes are an underestimated and probably important component of coastal marine ecosystems that are usually overlooked by standard methods for collecting and studying benthic fishes. Studies focusing on cryptobenthic fishes have been rare and all have been based on samples taken during daytime. The present study tested the difference in epibenthic and cryptobenthic fish composition, diel differences in cryptobenthic fish assemblage and diel shifts of infralittoral fish species between hidden and open bottom spaces. It also looked for the significant habitat variables structuring the cryptobenthic fish assemblage. The daylight, sunset and night samples of epibenthic and cryptobenthic fish assemblages were collected from 78 squares of 1 m2 shallow water plots (0.5 to 3 m) on Brač island in the eastern Adriatic. The study recorded 27 species, among which the family Gobiidae dominated fish diversity with 14 species. Cryptobenthic specimens highly outnumbered epibenthic specimens, with a ratio of 7.5 to 1. Species composition of cryptobenthic and epibenthic fish assemblages differed significantly. No diel variation in composition, species richness or abundance of the cryptobenthic fish assemblage was detected. Occurrence frequencies in hidden and open bottom spaces of ambivalent species did not change significantly between times of day, so no dial switches between open and hidden places were apparent. In combination, these results suggest that the cryptobenthic fish assemblage has diel stability and is mostly composed of permanent inhabitants of hidden spaces with domination of miniature gobies. Significant habitat variables for species occurrence were the presence of multiple layers, bottom inclination and the presence of cobbles, while depth also had a large but not significant effect.

ACS Style

Igor Glavičić; Marcelo Kovačić; Alen Soldo; Ulrich Schliewen. A quantitative assessment of the diel influence on the cryptobenthic fish assemblage of the shallow Mediterranean infralittoral zone. Scientia Marina 2020, 84, 1 .

AMA Style

Igor Glavičić, Marcelo Kovačić, Alen Soldo, Ulrich Schliewen. A quantitative assessment of the diel influence on the cryptobenthic fish assemblage of the shallow Mediterranean infralittoral zone. Scientia Marina. 2020; 84 (1):1.

Chicago/Turabian Style

Igor Glavičić; Marcelo Kovačić; Alen Soldo; Ulrich Schliewen. 2020. "A quantitative assessment of the diel influence on the cryptobenthic fish assemblage of the shallow Mediterranean infralittoral zone." Scientia Marina 84, no. 1: 1.

Journal article
Published: 27 December 2019 in Acta Adriatica
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Lesueurigobius friesii was collected in Eidsfjorden, Sognefjorden, Norway, extending its known distribution range north as the new northernmost locality of this species. Globally, the northernmost presence of gobies is along the coast of Norway. Their diversity along the Norwegian coast showed an evident latitude gradient of gobiid diversity with a clear decrease from south to north. The significant regression structural change was found at the 63/64° N latitude band followed by a 36.4% decrease in gobiid species diversity. The species traits of gobiids north of the regression breaking point and those restricted to the south of it were compared. The only significantly more frequent characteristic of species passing north of the regression breaking point is the large depth range that reach down to the shelf break. All species present north of the point, except Thorogobius ephippiatus (that barely passes it) belong to Oxudercinae (i.e. to Pomatoschistuslineage of that subfamily).

ACS Style

Marcelo Kovačić; Rudolf Svensen. Northern extension of Lesueurigobius friesii (Malm, 1874) (Pisces: Gobiidae) distribution and the gobiid diversity decline along the Norwegian coast. Acta Adriatica 2019, 60, 147 -156.

AMA Style

Marcelo Kovačić, Rudolf Svensen. Northern extension of Lesueurigobius friesii (Malm, 1874) (Pisces: Gobiidae) distribution and the gobiid diversity decline along the Norwegian coast. Acta Adriatica. 2019; 60 (2):147-156.

Chicago/Turabian Style

Marcelo Kovačić; Rudolf Svensen. 2019. "Northern extension of Lesueurigobius friesii (Malm, 1874) (Pisces: Gobiidae) distribution and the gobiid diversity decline along the Norwegian coast." Acta Adriatica 60, no. 2: 147-156.

Journal article
Published: 09 December 2019 in Zootaxa
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A new goby species, Lebetus patzneri sp. nov. (Teleostei: Gobiidae) is described based on two preserved and two photographed specimens collected from two circalittoral bottom locations off the Balearic Islands, western Mediterranean. It differs from the one or both other Lebetus species by each of following characters: P 15–16; D2 segmented fin rays 7–8; snout slightly larger than eye diameter; eyes small, 23.3–25.5% of head length; body height low, body depth at pelvic fin 14.9–16.4%; lips thick, giving mouth a broad, almost rectangular shape in dorsal and lateral view; minute black “pepper” spots on predorsal area, dorsum, flanks and caudal peduncle, visible in fresh and preserved specimens; the most of the caudal peduncle similarly colored as the lateral area below D2, reddish or brown. We further report first records of Lebetus guilleti (Le Danois, 1913) from Norway and the Balearic Islands, and observations of its conspicuous courtship behavior (“D1 waving”), and coloration including the first description of its red fluorescence pattern.

ACS Style

Ulrich K. Schliewen; Marcelo Kovačić; Alexander F. Cerwenka; Rudolf Svensen; F. (Francesc) Ordines. Lebetus patzneri (Teleostei: Gobiidae), a new goby species from the Balearic Islands, western Mediterranean, with first records of Lebetus guilleti (Le Danois, 1913) from this area and Norway, and with notes on its biology. Zootaxa 2019, 4706, 231 -254.

AMA Style

Ulrich K. Schliewen, Marcelo Kovačić, Alexander F. Cerwenka, Rudolf Svensen, F. (Francesc) Ordines. Lebetus patzneri (Teleostei: Gobiidae), a new goby species from the Balearic Islands, western Mediterranean, with first records of Lebetus guilleti (Le Danois, 1913) from this area and Norway, and with notes on its biology. Zootaxa. 2019; 4706 (2):231-254.

Chicago/Turabian Style

Ulrich K. Schliewen; Marcelo Kovačić; Alexander F. Cerwenka; Rudolf Svensen; F. (Francesc) Ordines. 2019. "Lebetus patzneri (Teleostei: Gobiidae), a new goby species from the Balearic Islands, western Mediterranean, with first records of Lebetus guilleti (Le Danois, 1913) from this area and Norway, and with notes on its biology." Zootaxa 4706, no. 2: 231-254.

Original article
Published: 18 November 2019 in Journal of Applied Ichthyology
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The recently described Mediterranean goby species, Buenia massutii, was found at the deep shelf in the Gulf of Cádiz (north‐eastern Atlantic Ocean). The identification of the specimen was performed using morphological analysis and molecular techniques; including the sequencing of the mitochondrial genes Cytochrome c Oxidase I (DNA barcode) and Cytochrome b. The sequences for Cytochrome b is presented for the first time for B. massutii. The limited migration capability and behaviour of small benthic gobies, the short lifespan of Buenia species, and the oceanographic characteristics of the Strait of Gibraltar suggest the existence of an independent Atlantic population of B. massutii.

ACS Style

Francesc Ordines; Sergio Ramírez‐Amaro; Candelaria Burgos; Jorge Baro; Marcelo Kovačić; Ignacio Sobrino. First record of Buenia massutii Kovačić, Ordines, & Schliewen, 2017 in the Atlantic Ocean based on molecular and morphological evidences. Journal of Applied Ichthyology 2019, 36, 85 -89.

AMA Style

Francesc Ordines, Sergio Ramírez‐Amaro, Candelaria Burgos, Jorge Baro, Marcelo Kovačić, Ignacio Sobrino. First record of Buenia massutii Kovačić, Ordines, & Schliewen, 2017 in the Atlantic Ocean based on molecular and morphological evidences. Journal of Applied Ichthyology. 2019; 36 (1):85-89.

Chicago/Turabian Style

Francesc Ordines; Sergio Ramírez‐Amaro; Candelaria Burgos; Jorge Baro; Marcelo Kovačić; Ignacio Sobrino. 2019. "First record of Buenia massutii Kovačić, Ordines, & Schliewen, 2017 in the Atlantic Ocean based on molecular and morphological evidences." Journal of Applied Ichthyology 36, no. 1: 85-89.

Journal article
Published: 15 September 2019 in Acta Ichthyologica et Piscatoria
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ACS Style

F. (Francesc) Ordines; Marcelo Kovačić; Miguel Vivas; Cristina García-Ruiz; Beatriz Guijarro. Westernmost Mediterranean records of three gobiid species (Actinopterygii: Perciformes: Gobiidae). Acta Ichthyologica et Piscatoria 2019, 49, 275 -282.

AMA Style

F. (Francesc) Ordines, Marcelo Kovačić, Miguel Vivas, Cristina García-Ruiz, Beatriz Guijarro. Westernmost Mediterranean records of three gobiid species (Actinopterygii: Perciformes: Gobiidae). Acta Ichthyologica et Piscatoria. 2019; 49 (3):275-282.

Chicago/Turabian Style

F. (Francesc) Ordines; Marcelo Kovačić; Miguel Vivas; Cristina García-Ruiz; Beatriz Guijarro. 2019. "Westernmost Mediterranean records of three gobiid species (Actinopterygii: Perciformes: Gobiidae)." Acta Ichthyologica et Piscatoria 49, no. 3: 275-282.

Journal article
Published: 06 August 2019 in Zootaxa
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A new genus and species of goby, Gymnesigobius medits sp. nov., is described from the western Mediterranean slope bottoms at the Balearic Islands. The new goby belongs to the Gobius-lineage (Gobiinae). Examination of the single known specimen exhibits a unique combination of morphological characters which could not be fitted to any known genus. Gymnesigobius gen. nov. is morphologically distinguished from all other genera in the Gobius-lineage by the following combination of characters: chin without fold or barbels; mouth terminal with anterior tip above horizontal level of lower eye edge; predorsal area and first dorsal fin base naked; pelvic fin anterior membrane well developed; head with anterior oculoscapular and preopercular canals, posterior oculoscapular canal absent; pores of head canals enlarged, e.g. pores α and ρ larger or of about the same size as interspaces to pore β; six transverse suborbital rows of sensory papillae, four continuous suborbital rows in front of row b, fifth row divided in three parts but in front of row b, sixth row just as superior part above row b and below pore α; longitudinal suborbital row b barely reaching forward to the vertical from posterior edge of eye. A full description of the new genus and species is provided. The new species, collected at a depth of between 344 and 364 m (mean depth of 354 m), is one of very few gobiid species found at bathyal depths.

ACS Style

Marcelo Kovačić; F. (Francesc) Ordines; Sergio Ramirez-Amaro; Ulrich K. Schliewen. Gymnesigobius medits (Teleostei: Gobiidae), a new gobiid genus and species from the western Mediterranean slope bottoms. Zootaxa 2019, 4651, 513 -530.

AMA Style

Marcelo Kovačić, F. (Francesc) Ordines, Sergio Ramirez-Amaro, Ulrich K. Schliewen. Gymnesigobius medits (Teleostei: Gobiidae), a new gobiid genus and species from the western Mediterranean slope bottoms. Zootaxa. 2019; 4651 (3):513-530.

Chicago/Turabian Style

Marcelo Kovačić; F. (Francesc) Ordines; Sergio Ramirez-Amaro; Ulrich K. Schliewen. 2019. "Gymnesigobius medits (Teleostei: Gobiidae), a new gobiid genus and species from the western Mediterranean slope bottoms." Zootaxa 4651, no. 3: 513-530.

Journal article
Published: 15 June 2019 in Acta Ichthyologica et Piscatoria
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ACS Style

Marcelo Kovačić; Igor Glavičić. The first Adriatic finding of Speleogobius llorisi (Actinopterygii: Gobiiformes: Gobiidae). Acta Ichthyologica et Piscatoria 2019, 49, 181 -184.

AMA Style

Marcelo Kovačić, Igor Glavičić. The first Adriatic finding of Speleogobius llorisi (Actinopterygii: Gobiiformes: Gobiidae). Acta Ichthyologica et Piscatoria. 2019; 49 (2):181-184.

Chicago/Turabian Style

Marcelo Kovačić; Igor Glavičić. 2019. "The first Adriatic finding of Speleogobius llorisi (Actinopterygii: Gobiiformes: Gobiidae)." Acta Ichthyologica et Piscatoria 49, no. 2: 181-184.

Short communication
Published: 31 May 2019 in Molecular Phylogenetics and Evolution
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We report a hitherto unknown radiation within the clingfishes (Gobiesocidae), discovered in one of the best-studied marine biomes, the Mediterranean Sea. The monotypic genus Gouania is a Mediterranean endemic inhabiting the interstices of gravel beaches. Using geometric morphometric analyses, we identified two distinct morphotypes (characterized by a slender and a stout body shape, respectively) among Gouania willdenowi sampled from the three major Mediterranean basins (Eastern, Western and Adriatic). Slender and stout G. willdenowi occurred sympatrically in the Adriatic and the Eastern Mediterranean basins. Only the stout morphotype was found in the Western Mediterranean. Morphotypes were further distinguished by relative eye size and number of vertebrae. Based on mitochondrial and nuclear gene sequences, five highly divergent clades (COI-based K2P distances of 8 to 15%) were identified. The clades represented, respectively, stout Adriatic, slender Adriatic, stout Eastern, slender Eastern and stout Western Mediterranean Gouania. This suggests that the genus Gouania comprises at least five different species. Phylogenetic relationships among clades furthermore imply repeated evolution of convergent morphotypes. The onset of the Gouania radiation was dated to 3.23 (95% HPD 2.08 – 5.90) mya, and may therefore have overlapped with or followed the Messinian salinity crisis.

ACS Style

Maximilian Wagner; Sandra Bračun; Gerhard Skofitsch; Marcelo Kovačić; Stamatis Zogaris; Samuel Iglésias; Kristina M. Sefc; Stephan Koblmüller. Diversification in gravel beaches: A radiation of interstitial clingfish (Gouania, Gobiesocidae) in the Mediterranean Sea. Molecular Phylogenetics and Evolution 2019, 139, 106525 .

AMA Style

Maximilian Wagner, Sandra Bračun, Gerhard Skofitsch, Marcelo Kovačić, Stamatis Zogaris, Samuel Iglésias, Kristina M. Sefc, Stephan Koblmüller. Diversification in gravel beaches: A radiation of interstitial clingfish (Gouania, Gobiesocidae) in the Mediterranean Sea. Molecular Phylogenetics and Evolution. 2019; 139 ():106525.

Chicago/Turabian Style

Maximilian Wagner; Sandra Bračun; Gerhard Skofitsch; Marcelo Kovačić; Stamatis Zogaris; Samuel Iglésias; Kristina M. Sefc; Stephan Koblmüller. 2019. "Diversification in gravel beaches: A radiation of interstitial clingfish (Gouania, Gobiesocidae) in the Mediterranean Sea." Molecular Phylogenetics and Evolution 139, no. : 106525.

Journal article
Published: 21 May 2019 in Zootaxa
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Two new species of the gobiid genus Hetereleotris, H. aurantiaca sp. nov. and H. semisquamata sp. nov., are described from the Red Sea, the former from Saudi Arabia at Jeddah from the cave at depth of 14-16 m, and the latter from the southern Egypt from reef flat. Hetereleotris aurantiaca sp. nov. is distinguished from its congeners by having dorsal-fin rays VI + I,10; anal-fin rays I,9; pectoral-fin rays 14, all rays branched; pelvic-fin rays I,5, the fin separated and without frenum, 5th ray unbranched; anterior nostril with a long tube without process from the rim, posterior nostril a pore with erected rim; no tentacle above eye; posterior angle of jaws extending posteriorly to below posterior edge of pupil; no opercular spine; no mental frenum; pelvic fins longer than pectoral fins; squamation reduced to a few scales on caudal peduncle at caudal-fin base; no head canals; by presence, size and pattern of suborbital rows of sensory papillae; and orange head and yellowish orange body with five faint brown bars. Hetereleotris semisquamata sp. nov. is distinctive among its congeners by unique scale pattern (scales cycloid, the squamation reduced, tapering from caudal-fin base along lateral midline towards pectoral fin where nearly reaching its base) and by coloration (head and body whitish, with brown line from eye to end of upper lip, dark brown band across interorbital area and continuing obliquely from eye to corner of opercle, broad dark brown band below first dorsal fin continuing into fin, and moderately broad dark brown bar on caudal-fin base). Furthermore, it is characterized in having dorsal-fin rays VI + I,11, anal-fin rays I,10, pectoral-fin rays 16, and absence of head canals. In addition to descriptions of two species, a key to all species of Hetereleotris is provided. Hetereleotris psammophila is reported outside the Gulf of Aqaba for the first time.

ACS Style

Marcelo Kovačić; Sergey V. Bogorodsky; Ahmad O. Mal. Two new species of Hetereleotris (Perciformes: Gobiidae) from the Red Sea. Zootaxa 2019, 4608, 501 -516.

AMA Style

Marcelo Kovačić, Sergey V. Bogorodsky, Ahmad O. Mal. Two new species of Hetereleotris (Perciformes: Gobiidae) from the Red Sea. Zootaxa. 2019; 4608 (3):501-516.

Chicago/Turabian Style

Marcelo Kovačić; Sergey V. Bogorodsky; Ahmad O. Mal. 2019. "Two new species of Hetereleotris (Perciformes: Gobiidae) from the Red Sea." Zootaxa 4608, no. 3: 501-516.

Journal article
Published: 17 May 2019 in Mediterranean Marine Science
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Twelve new first records of species from two families are added to the list of known marine fishes from Malta based on laboratory study of material collected during fieldwork over a period of more than twenty years. The three new records of clingfishes (Apletodon incognitus, Gouania wildenowi, Opeatogenys gracilis) double the known diversity of Gobiesocidae for the area; the nine new records of gobies (Chromogobius zebratus, Gobius ater, Gobius fallax, Gobius gasteveni, Gobius roulei, Lebetus guilleti, Odondebuenia balearica, Speleogobius llorisi, Vanneaugobius dollfusi) increase by almost a half the known diversity of Maltese Gobiidae. The most enigmatic species reported is G. ater, for which the most recent published record is two decades old and based on specimens that had actually been collected back in the 19th century. Checklists of the Gobiesocidae and Gobiidae of Maltese waters are presented. Biodiversity and biogeographic implications of the present findings are discussed.

ACS Style

Marcelo Kovačić; Patrick J. Schembri. Twelve new records of gobies and clingfishes (Pisces: Teleostei) significantly increase small benthic fish diversity of Maltese waters. Mediterranean Marine Science 2019, 20, 287 -296.

AMA Style

Marcelo Kovačić, Patrick J. Schembri. Twelve new records of gobies and clingfishes (Pisces: Teleostei) significantly increase small benthic fish diversity of Maltese waters. Mediterranean Marine Science. 2019; 20 (2):287-296.

Chicago/Turabian Style

Marcelo Kovačić; Patrick J. Schembri. 2019. "Twelve new records of gobies and clingfishes (Pisces: Teleostei) significantly increase small benthic fish diversity of Maltese waters." Mediterranean Marine Science 20, no. 2: 287-296.