A new mosasaur, Russellosaurus coheni, is described from the Collignoniceras woollgari Zone (lower Middle Turonian) at Cedar Hill, Dallas County, Texas. At approximately 92 Ma, it is the oldest well-preserved mosasaur skull from North America. It possesses characters diagnostic of Plioplatecarpinae but retains numerous plesiomorphies as well. Phylogenetic analysis indicates a close relationship with Yaguarasaurus columbianus, and these two, together with Tethysaurus nopcsai, form a clade that occupies a position basal to the divergence of the subfamilies Tylosaurinae and Plioplatecarpinae. Russellosaurus coheni is proposed as the nominal taxon of a new mosasaur clade, parafamily taxon novum Russellosaurina, which includes Plioplatecarpinae, Tylosaurinae, their common ancestor and all descendants. Tethysaurus retains a plesiopedal limb and girdle morphology, and along with Russellosaurus and Yaguarasaurus, cranial plesiomorphies. Dallasaurus turneri, a temporally and geographically sympatric plesiopedal mosasaur, occupies a basal position within Mosasaurinae. This phyletic arrangement confirms that marine adaptations, such as development of paddle-like limbs, occurred independently in at least two lineages of mosasaurs, once within Mosasaurinae and once within Russellosaurina.
Doi:10.4202/app.2008.0063 A new Late Cretaceous turtle, Angolachelys mbaxi gen. et sp. nov., from the Turonian (90 Mya) of Angola, represents the oldest eucryptodire from Africa. Phylogenetic analysis recovers Angolachelys mbaxi as the sister taxon of Sandownia harrisi from the Aptian of Isle of Wight, England. An unnamed turtle from the Albian Glen Rose Formation of Texas (USA) and the Kimmeridgian turtle Solnhofia parsonsi (Germany), are successively more distant sister taxa. Bootstrap analysis suggests those four taxa together form a previously unrecognized monophyletic clade of marine turtles, herein named Angolachelonia clade nov., supported by the following synapomorphies: mandibular articulation of quadrate aligned with or posterior to the occiput, and basisphenoid not visible or visibility greatly reduced in ventral view. Basal eucryptodires and angolachelonians originated in the northern hemisphere, thus Angolachelys represents one of the firs
25 September 2012 in Earth and Environmental Science Transactions of the Royal Society of Edinburgh
Upper Cretaceous marine rocks of the Big Bend Region of trans-Pecos Texas preserve a number of marine-adapted mosasauroids. At least three unnamed taxa of basal mosasauroids are represented by remains from shaly limestones in the middle Turonian portion of the Boquillas Formation. These occur along with remains of larger derived mosasaurs referable to Russellosaurina and an undescribed tylosaurine. Derived mosasaurs from the middle to late Coniacian include the first report of Tylosaurus kansasensis outside of Kansas, T. nepaeolicus, Platecarpus planifrons, and Platecarpus aff. P. planifrons. Clidastes liodontus is found in the latest Coniacian or early Santonian part of the Pen Formation. An undescribed species of Ectenosaurus, Clidastes sp. and an indeterminate plioplatecarpine occurs in the middle Santonian to early Campanian interval of the Pen Formation. The mosasaur fauna from the Big Bend region is quite similar to that from the Smoky Hill Chalk of Kansas, a thousand kilometres to the north. We refine the position of the Cenomanian–Turonian boundary within the Ernst Member of the Boquillas Formation, based on ammonite faunas. We also corroborate previous interpretations describing the time-transgressive nature of the onset of deposition of the Pen Formation based on a west-to-east descending level of the Inoceramus (Cremnoceramus) undulatoplicatus FAD.
Here, we describe the first pterosaur remains from Angola, an assemblage of fourteen bones from the Lower Maastrichtian marine deposits of Bentiaba, Namibe Province. One new species is introduced, Epapatelo otyikokolo, gen. et sp. nov., which comprises an articulated partial left humerus and ulna as well as an articulated left ulna and radius (from a second individual). Phylogenetic analysis confirms a non-nyctosaurid pteranodontian attribution for this new taxon and supports a new apomorphy-based clade, Aponyctosauria, which is here defined. Late Cretaceous pteranodontians are rare in Sub-Saharan Africa and throughout the Southern Hemisphere. Preliminary histological analysis also reveals a likely sub-adult age for one of the specimens. This fossil assemblage provides a first glimpse of Angolan pterosaur paleobiodiversity providing further insight into the Gondwanan ecosystems of the Upper Cretaceous.