We developed ecologically based climate‐fire projections for the western U.S. Using a finer ecological classification and fire‐relevant climate predictors, we created statistical models linking climate and wildfire area burned for ecosections, which are geographic delineations based on biophysical variables. The results indicate a gradient from purely fuel‐limited (antecedent positive water balance anomalies or negative energy balance anomalies) to purely flammability‐limited (negative water balance anomalies or positive energy balance anomalies) fire regimes across ecosections. Although there are other influences (such as human ignitions and management) on fire occurrence and area burned, seasonal climate significantly explains interannual fire area burned. Differences in the role of climate across ecosections are not random, and the relative dominance of climate predictors allows objective classification of ecosection climate‐fire relationships. Expected future trends in area burned range from massive increases, primarily in flammability limited systems near the middle of the water balance deficit distribution, to substantial decreases, in fuel‐limited non‐forested systems. We predict increasing area burned in most flammability‐limited systems but predict decreasing area burned in primarily fuel‐limited systems with a flammability‐limited (“hybrid”) component. Compared to 2030‐2059 (2040s), projected area burned for 2070‐2099 (2080s) increases much more in the flammability and flammability dominated hybrid systems than those with equal control, and continues to decrease in fuel‐limited hybrid systems. Exceedance probabilities for historical 95^th percentile fire years are larger in exclusively flammability‐limited ecosections than in those with fuel controls. Filtering the projected results using a fire‐rotation constraint minimizes over‐projection due to static vegetation assumptions, making projections more conservative.

Across the Upper Missouri River Basin, the recent drought of 2000 to 2010, known as the “turn-of-the-century drought,” was likely more severe than any in the instrumental record including the Dust Bowl drought. However, until now, adequate proxy records needed to better understand this event with regard to long-term variability have been lacking. Here we examine 1,200 y of streamflow from a network of 17 new tree-ring–based reconstructions for gages across the upper Missouri basin and an independent reconstruction of warm-season regional temperature in order to place the recent drought in a long-term climate context. We find that temperature has increasingly influenced the severity of drought events by decreasing runoff efficiency in the basin since the late 20th century (1980s) onward. The occurrence of extreme heat, higher evapotranspiration, and associated low-flow conditions across the basin has increased substantially over the 20th and 21st centuries, and recent warming aligns with increasing drought severities that rival or exceed any estimated over the last 12 centuries. Future warming is anticipated to cause increasingly severe droughts by enhancing water deficits that could prove challenging for water management. Significance Recent decades have seen droughts across multiple US river basins that are unprecedented over the last century and potentially longer. Understanding the drivers of drought in a long-term context requires extending instrumental data with paleoclimatic data. Here, a network of new millennial-length streamflow reconstructions and a regional temperature reconstruction from tree rings place 20th and early 21st century drought severity in the Upper Missouri River basin into a long-term context. Across the headwaters of the United States’ largest river basin, we estimated region-wide, decadal-scale drought severity during the “turn-of-the-century drought” ca. 2000 to 2010 was potentially unprecedented over the last millennium. Warming temperatures have likely increasingly influenced streamflow by decreasing runoff efficiency since at least the late 20th century.

Climatically driven changes in snow characteristics (snowfall, snowpack, and snowmelt) will affect hydrologic and ecological systems in Alaska over the coming century, yet there exist no projections of downscaled future snow pack metrics for the state of Alaska. We updated historical and projected snow day fraction (PSF, the fraction of days with precipitation falling as snow) from McAfee et al. We developed modeled snowfall equivalent (SFE) derived from the product of snow-day fraction (PSF) and existing gridded precipitation for Alaska from Scenarios Network for Alaska and Arctic Planning (SNAP). We validated the assumption that modeled SFE approximates historical decadally averaged snow water equivalent (SWE) observations from snowcourse and Snow Telemetry (SNOTEL) sites. We present analyses of future downscaled PSF and two new products, October–March SFE and ratio of snow fall equivalent to precipitation (SFE:P) based on bias-corrected statistically downscaled projections of Coupled Model Intercomparison Project 5 (CMIP5) Global Climate Model (GCM) temperature and precipitation for the state of Alaska. We analyzed mid-century (2040–2069) and late-century (2070–2099) changes in PSF, SFE, and SFE:P relative to historical (1970–1999) mean temperature and present results for Alaska climate divisions and 12-digit Hydrologic Unit Code (HUC12) watersheds. Overall, estimated historical the SFE is reasonably well related to the observed SWE, with correlations over 0.75 in all decades, and correlations exceeding 0.9 in the 1960s and 1970s. In absolute terms, SFE is generally biased low compared to the observed SWE. PSF and SFE:P decrease universally across Alaska under both Representative Concentration Pathway (RCP) 4.5 and RCP 8.5 emissions scenarios, with the smallest changes for RCP 4.5 in 2040–2069 and the largest for RCP 8.5 in 2070–2099. The timing and magnitude of maximum decreases in PSF vary considerably with regional average temperature, with the largest changes in months at the beginning and end of the snow season. Mean SFE changes vary widely among climate divisions, ranging from decreases between −17 and −58% for late twenty-first century in southeast, southcentral, west coast and southwest Alaska to increases up to 21% on the North Slope. SFE increases most at highest elevations and latitudes and decreases most in coastal southern Alaska. SFE:P ratios indicate a broad switch from snow-dominated to transitional annual hydrology across most of southern Alaska by mid-century, and from transitional to rain-dominated watersheds in low elevation parts of southeast Alaska by the late twenty-first century.

Under climate change, ecosystems are experiencing novel drought regimes, often in combination with stressors that reduce resilience and amplify drought’s impacts. Consequently, drought appears increasingly likely to push systems beyond important physiological and ecological thresholds, resulting in substantial changes in ecosystem characteristics persisting long after drought ends (i.e., ecological transformation). In the present article, we clarify how drought can lead to transformation across a wide variety of ecosystems including forests, woodlands, and grasslands. Specifically, we describe how climate change alters drought regimes and how this translates to impacts on plant population growth, either directly or through drought's interactions with factors such as land management, biotic interactions, and other disturbances. We emphasize how interactions among mechanisms can inhibit postdrought recovery and can shift trajectories toward alternate states. Providing a holistic picture of how drought initiates long-term change supports the development of risk assessments, predictive models, and management strategies, enhancing preparedness for a complex and growing challenge.