In the Western United States, volcanic spring-fed rivers are anticipated to become increasingly more important for salmonids and other native fishes, as these rivers will retain coldwater habitats as the climate warms. Despite this, little is known about the hydro-biogeochemical interactions within these ecosystems. A review of existing literature on spring-fed rivers, coupled with a decade of research on volcanic spring-fed rivers of northern California, finds that these systems are exceptionally productive and exhibit stable environmental conditions. These unique conditions stem from hydrogeologic processes typical of young volcanic terrains. Aquatic macrophytes, common to some nutrient-rich spring-fed systems, play a disproportionate role in hydrologic and geomorphic processes by facilitating ecological interactions and velocity conditions that improve juvenile salmonid growth. We find that volcanic spring-fed rivers are also resilient to climate change, due not only to their ability to dampen water temperature changes through deep groundwater flow but also because of their nutrient-driven high ecosystem productivity, which may enable coldwater species to metabolically compensate for marginal increases in water temperature. Understanding the fundamental geomorphic and ecological differences between these rare ecosystems and their numerically dominant runoff rivers is essential for developing long-term conservation strategies for coldwater species under a rapidly changing climate.

Reservoir-based hydropower systems represent key interactions between water and energy systems and are being transformed under policy initiatives driven by increasing water and energy demand, the desire to reduce environmental impacts, and interacting effects of climate change. Such policies are often guided by complex system models, whereby divergence in system representations can potentially translate to incompatible planning outcomes, thereby undermining any planning that may rely on them. We review different approaches and assumptions in hydropower representation in water and energy systems. While the models and issues are relevant globally, the review focuses on applications in California given its extensive development of energy and water models for policy planning, but discusses the extent to which these observations apply to other regions. Structurally, both water-driven and energy-driven management models are similar. However, in energy models, hydropower is often represented as a single-priority output. Water management models typically allocate water for competing priorities, which are generally uninformed by dynamic electricity load demand, and often result in a lower priority for hydropower. In water models, constraints are increasingly resolved for non-energy components (e.g. inflow hydrology and non-energy water demand); few analogues exist for energy models. These limitations may result in inadequate representations of each respective sector, and vastly different planning outcomes for the same facilities between the two different sectors. These divergent modeling approaches manifest themselves in California where poorly reconciled outcomes may affect decisions in hydropower licensing, electricity grid flexibility and decarbonization, and planning for environmental water. Fully integrated water-energy models are computationally intensive and specific to certain regions, but better representation of each domain in respective efforts would help reconcile divergences in planning and management efforts related to hydropower across energy and water systems.

Environmental flows, or the practice of allocating water in river systems for ecological purposes, is a leading strategy for conserving aquatic species and improving river health. However, consideration of surface-groundwater connectivity is seldom addressed in environmental flow development due to a lack of methodologies that account for groundwater contributions to instream flow. Groundwater-influenced streams have been identified as key refugia for native biota under a rapidly changing climate. These ecosystems are anticipated to be more resistant to climate change because groundwater input buffers the adverse effects of low flows and high temperatures, particularly in the dry season. Less understood, however, is the relative contribution of groundwater inputs to streamflow and how these surface-groundwater water interactions should be accounted for in environmental flow assessments and management actions. In order to assess ecological flow needs in groundwater-influenced streams, we applied the California Environmental Flows Framework (CEFF) in two river systems in California, United States. The Little Shasta River and the lower Cosumnes River are representative of many groundwater-influenced streams throughout the semi-arid western United States. Historically, perennial streamflow once sustained diverse native aquatic species in these ecosystems, but water withdrawals for irrigated agriculture has resulted in periodic stream dewatering. We found CEFF was useful in quantifying ecological flow needs for seasonal components of the flow regime that support ecosystem functionality. In particular, CEFF offered flexibility to incorporate information on the seasonal and spatial dimensions of groundwater influences in the development of ecological flow targets. The focus on ecosystem functions in CEFF, and ability to account for groundwater influences on those functions, creates opportunities for integrated surface-groundwater management strategies that support the recovery and protection of streamflows in groundwater-influenced streams.

Stream temperature science and management is rapidly shifting from single-metric driven approaches to multi-metric, thermal regime characterizations of streamscapes. Given considerable investments in recovery of cold-water fisheries (e.g., Pacific salmon and other declining native species), understanding where cold water is likely to persist, and how cold-water thermal regimes vary, is critical for conservation. California’s unique position at the southern end of cold-water ecosystems in the northern hemisphere, variable geography and hydrology, and extensive flow regulation requires a systematic approach to thermal regime classification. We used publicly available, long-term (> 8 years) stream temperature data from 77 sites across California to model their thermal regimes, calculate three temperature metrics, and use the metrics to classify each regime with an agglomerative nesting algorithm. Then, we assessed the variation in each class and considered underlying physical or anthropogenic factors that could explain differences between classes. Finally, we considered how different classes might fit existing criteria for cool- or cold-water thermal regimes, and how those differences complicate efforts to manage stream temperature through regulation. Our results demonstrate that cool- and cold-water thermal regimes vary spatially across California. Several salient findings emerge from this study. Groundwater-dominated streams are a ubiquitous, but as yet, poorly explored class of thermal regimes. Further, flow regulation below dams imposes serial discontinuities, including artificial thermal regimes on downstream ecosystems. Finally, and contrary to what is often assumed, California reservoirs do not contain sufficient cold-water storage to replicate desirable, reach-scale thermal regimes. While barriers to cold-water conservation are considerable and the trajectory of cold-water species towards extinction is dire, protecting reaches that demonstrate resilience to climate warming remains worthwhile.

Maximum water temperatures in streams throughout the western USA typically occur in late summer and early autumn, coinciding with low stream flow. However, in the spring-fed Big Springs Creek in northern California, where constant-temperature groundwater springs provide relatively stable stream flow throughout the year, peak water temperatures and maximum diurnal variability occur in spring. We attribute this anomaly to the riverine canopy provided by emergent aquatic macrophytes (e.g., Polygonum hydropiperoides and Nasturtium officinale), which mimics the shade function of a riparian canopy. Macrophyte biomass increased 264% between January and August 2011. This increase coincided with a 111% reduction in flow velocity and a 53% increase in stream depth. Solar radiation was reduced by an average of 88% in patches of macrophytes that covered ~50% of the water surface during the summer. Decreased solar radiation reduced rates of stream heating, maximum temperatures, and temperature variability. We tested the riverine canopy hypothesis analytically based on a 2-dimensional hydrodynamic and water-temperature model. The model predicted that emergent aquatic macrophytes reduce maximum water temperatures by an average of 5.1°C (p < 0.001) during late summer, when water temperatures in northern California streams typically increase. Our study shows the influence of a riverine canopy on naturally occurring temporal patterns of water temperature in a spring-fed stream. Our results could inform basin-scale management or regulatory strategies to address water-temperature conditions.

Environmental flow management in watersheds with multi-objective reservoirs is often presented as an additional constraint to an already strained and over-allocated stream system. Nevertheless, environmental flow legislation and regulatory policies are increasingly being developed and implemented globally. In California, USA, recent legislative and regulatory policies place environmental flows at the forefront of the state’s water management objectives; however, the increased reliance on hydropower to support climate change mitigation goals may complicate efforts on both issues. This study modelled alternative environmental flow strategies in the major tributaries to the San Joaquin River in California. Strategies included detailed water management rules for hydropower production, flood control, and water deliveries, and three methodological approaches to environmental flow releases: minimum instream flows (“baseline”) year-round, 40% of full natural flow (FNF) during the spring runoff season and minimum releases the remainder of the year, and functional flows year-round. Results show that environmental flow strategies affect downstream flow releases in each of the San Joaquin’s four sub-basins differently depending on infrastructure capacity, water management objectives, and hydrologic year types. While hydropower production was comparable or declined in the Stanislaus, Tuolumne, and Merced basins, functional flow and 40% FNF strategies increased hydropower production in the Upper San Joaquin basin by 11%. Uncontrolled spill of high flow events decreased when high flow releases were based on hydrologic cues rather than exclusively on flood storage capacity. Water deliveries were reduced in all years regardless of environmental flow strategy. The 40% FNF and functional flow strategies both increased water released to the river relative to baseline, but in different ways. The functional flow strategy allocated water in a holistic approach that enhanced ecological functions in all years, but particularly in moderate and wet years. In contrast, the 40% FNF strategy provided increased flows relative to baseline and some ecological benefit in dry years, but less ecological benefit in other years. This study shows that alternative environmental flow strategies will have different and important trade-offs for integrated water management, and may mutually benefit seemingly conflicting objectives.