The area of foliage absorbing solar radiation is often expressed as leaf area index (LAI). In this study, specific leaf area (SLA), leaf area at tree level (LA) and LAI at stand level were measured on eight experimental plots of Norway spruce (Picea abies) and Scots pine (Pinus sylvestris), together with tree and stand measurements from biomass harvest. Both projected area and half total surface area were measured, and a model of the relationship between the units was constructed. SLA was larger for pine (61.62 cm2 g−1) compared to spruce (50.2 cm2 g−1) and showed a trend of decreasing higher up in the crown. Leaf area was significantly higher for Norway spruce compared to Scots pine on both tree and stand level. Models were constructed using diameter at breast height, tree height and stand basal area to estimate LA. The models were general site-independent models that can be used for easy estimations of single tree leaf area. Indirect measurement of LAI (LAIe) was shown to underestimate LAI with on average 30–73% depending on species and measurement technique. Using the extensive data collected, conversion models were constructed for estimating LAI from LAIe together with basal area, stem number and stand height. These species-specific conversion models will allow for more accurate estimations of LAI that can be used in mechanistic models of forest growth and for future estimates of LAI from remote sensing data.
2 January 2022 in Scandinavian Journal of Forest Research
When modelling forest growth, capturing the effects of climate change is needed for reliable long-term predictions and management choices. This remains a challenge because commonly used mensurational forest growth and yield models, relying on inventory data, cannot account for climate change effects. We developed hybrid physiological/mensurational basal area growth and yield models, which combine physiological response to climatic conditions and empirical relations. We included climate and site effects by replacing time with light sums of photosynthetically active radiation and modifying the latter with monthly soil water, vapour pressure deficit, temperature, and frost days. When parameterised with permanent sample plot data for Scots pine and Norway spruce across Sweden, the hybrid models could reproduce observations well, although with no increase in precision compared with time-based mensurational models. When considering different climate scenarios, a significant impact on productivity from climate change emerged. For example, a 2 °C warming enhanced Scots pine production by up to 14% in regions where temperatures were originally cooler and soil water deficit was low (i.e. northwest Sweden), but depressed it, up to 9%, elsewhere. Hence, climate-sensitive models that take local variations into account are necessary for accurate predictions and sustainable forest management.
11 February 2021 in Scandinavian Journal of Forest Research
Admixtures of birch in Norway spruce plantations are being promoted as a means to increase habitat and species diversity. The implications of this mixture were analysed with regional survey data from southern Sweden. Permanent sample plots from the Swedish National Forest Inventory (NFI), with Norway spruce and admixture of birch, were used to describe the temporal trends in the admixture, regarding species composition and competitive strength. Observations from thinned plots show a higher harvest removal in birch (35%) than for Norway spruce (19%). Observations without thinnings in the period before measurement showed that individual birch tree growth was lower compared to Norway spruce and it decreased even more with increasing stand age and competition. In addition, a complementary field survey, with multiple distributed sample plots in each stand, was used to detect within-stand variation of species composition and density. Although within-stand heterogeneity was larger in mixed stands in terms of species composition, it was not different from Norway spruce monocultures in terms of stand density. These two surveys show that the admixture of birch, for several reasons, decreases over stand age and although birch increases tree species diversity, it does not necessary imply a change in density.
Definitions of continuous cover forestry (CCF) vary among countries, and are often a political compromise. We offer a common definition of CCF for this book, which can be found in a text box below. The three silvicultural systems included in CCF are described briefly. Conversion to CCF will be an important activity in the near future, but approaches to and experiences of conversion to CCF are largely lacking in Fennoscandia. Methods need to be developed for how to assess the suitability of forest stands for CCF or conversion to CCF. Bad practices and experiences with selective cutting in Fennoscandia before 1950 have led to a loss of experience and willingness to apply CCF. Climate adaptation will make it necessary to modify CCF approaches in the region, especially given the limited number of shade-tolerant species.
In the context of continuous cover forestry (CCF), natural regeneration is the preferred form of regeneration, but it is a long-lasting and complex process. Shelter density has a large effect on the regeneration process and results. The selection system, particularly suited for shade-tolerant species like Norway spruce, relies on continuous regeneration and ingrowth into larger size classes. Regeneration and ingrowth rates vary significantly among stands, influenced by site and historical factors, with no clear relationship to current stand conditions. In the group system, edge trees influence regeneration by providing seeds, checking weed growth, and exerting competition. Regeneration in gaps is generally satisfactory for both Norway spruce and Scots pine. However, seedlings usually grow slower, especially close to the gap edges. The shelterwood system promotes regeneration through a successive, uniform opening of the canopy. Shelter trees provide seeds, and reduce seedling damage and competition from ground vegetation. On the other hand, the remaining overstorey shelter trees reduce seedling growth. Conversion to the selection system initiates regeneration in young stands, aiming for slow and steady regeneration. Given the rapid growth and crown closure in young stands, frequent manipulation of shelter density is essential during conversion, for example by opening small gaps.