Mikolaj Lula

Dr. Mikolaj Lula


Research Focus & Metrics

Academic Career Length

6Years (2021-Present)

(Calculated once Scilit publication data is synced: current year − year of first publication + 1)

Representative Publications

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6 April 2021 in Scandinavian Journal of Forest Research
Verified
Journal Article
Open access
MLMikolaj Lula
Mikolaj Lula
RTRenats Trubins
Renats Trubins
PEPer Magnus Ekö
Per Magnus Ekö
UJUlf Johansson
Ulf Johansson
UNUrban Nilsson
Urban Nilsson
Despite numerous studies there are still uncertainties regarding regeneration strategies that are optimal for productivity and profitability. Thus the aim of this study was to establish effects of three regeneration methods (planting, direct seeding and natural regeneration) on the production and profitability of Scots pine (Pinus sylvestris L.) stands in southern Sweden. Long-term stand development was simulated, with the StandWise application of the Heureka decision support system, starting from short-term regeneration outcomes observed in several field experiments at sites with relatively high productivity (H100 site indices, i.e. heights of dominant pines at 100 years: 27–30 m). Financial and production results of each approach were assessed in terms of Land Expectation Value (LEV) and Mean Annual Increment (MAI), respectively, across a whole rotation. Planting on clear-cuts with 1600–3265 seedlings per hectare resulted in the highest profitability and production, whereas high-density planting (10,000 seedlings per hectare) resulted in negative LEV. However, sensitivity analysis showed that the results depended on the interest rate. Retention of seed-trees incurred additional costs relative to single-operation clear felling. In contrast, retention of shelter-trees had good financial results (at 0% and 2.5% interest rate), although they depended on the site index and average tree size.

6 December 2024
Verified
Book Chapter
Open access
PDF file
MLMikolaj Lula
Mikolaj Lula
KHKjersti Holt Hanssen
Kjersti Holt Hanssen
MGMartin Goude
Martin Goude
HHHannu Hökkä
Hannu Hökkä
SVSauli Valkonen
Sauli Valkonen
AGAksel Granhus
Aksel Granhus
In the context of continuous cover forestry (CCF), natural regeneration is the preferred form of regeneration, but it is a long-lasting and complex process. Shelter density has a large effect on the regeneration process and results. The selection system, particularly suited for shade-tolerant species like Norway spruce, relies on continuous regeneration and ingrowth into larger size classes. Regeneration and ingrowth rates vary significantly among stands, influenced by site and historical factors, with no clear relationship to current stand conditions. In the group system, edge trees influence regeneration by providing seeds, checking weed growth, and exerting competition. Regeneration in gaps is generally satisfactory for both Norway spruce and Scots pine. However, seedlings usually grow slower, especially close to the gap edges. The shelterwood system promotes regeneration through a successive, uniform opening of the canopy. Shelter trees provide seeds, and reduce seedling damage and competition from ground vegetation. On the other hand, the remaining overstorey shelter trees reduce seedling growth. Conversion to the selection system initiates regeneration in young stands, aiming for slow and steady regeneration. Given the rapid growth and crown closure in young stands, frequent manipulation of shelter density is essential during conversion, for example by opening small gaps.

13 August 2026 in Forests
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Journal Article
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Ana Cristina GonçalvesAna Cristina Gonçalves
Ana Cristina Gonçalves
PSPeter Spathelf
Peter Spathelf
MLMikolaj Lula
Mikolaj Lula
TFTeresa Fonseca
Teresa Fonseca
Pines (genus Pinus) are a diverse group of conifers widely distributed across the Northern Hemisphere, comprising more than 110 species worldwide. In Europe, pines are a defining component of forests, occupying environments that range from dry Mediterranean lowlands to high-elevation alpine zones. They support a wide spectrum of ecosystem services, from timber and non-wood products to protection, biodiversity, and cultural values. In this review, we bring together ecological traits, silvicultural practices, and management considerations for ten European pine species. This review examines ten major European pine species, including Scots pine (Pinus sylvestris), maritime pine (P. pinaster), stone pine (P. pinea), Aleppo pine (P. halepensis), black pine (P. nigra), mountain pine (P. mugo), Swiss stone pine (P. cembra), Turkish pine (P. brutia), Bosnian pine (P. heldreichii), and Macedonian pine (P. peuce). Together, these species span a wide range of European environments, with Pinus sylvestris showing the broadest distribution. Despite the extensive literature on pine species, information on their ecological, silvicultural, and management attributes remains fragmented across disciplines. To bridge this gap, this review draws on an expert-curated core of scientific and technical literature complemented by a structured literature search adapted from the PRISMA framework, compiling within a single framework their main ecological and silvicultural characteristics, altitudinal range, growth patterns, stand diversity, and management practices and the principal goods and ecosystem services they provide. The comparative analysis highlighted distinct functional groups among European pines, illustrating contrasting adaptive strategies across environmental gradients, with altitudinal distributions ranging from sea level to 2400 m. Mediterranean species, such as P. halepensis, P. pinaster, and P. brutia, generally showed higher drought and fire adaptation, whereas mountain taxa such as P. cembra, P. mugo, and P. heldreichii displayed greater cold tolerance but increased vulnerability due to restricted climatic niches and slow regeneration. Broadly distributed species, such as P. sylvestris and P. nigra, exhibited intermediate ecological strategies, with responses strongly influenced by regional climatic conditions. By integrating evidence on distribution patterns, functional traits, growth patterns, and management practices, this review provides a comparative framework to support adaptive forest management and informed species selection, helping to safeguard forest functions and their associated benefits under ongoing climate change.

1 January 2026 in Silva Fennica
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Journal Article
Open access
PDF file
TNTherese Strömvall Nyberg
Therese Strömvall Nyberg
MLMikolaj Lula
Mikolaj Lula
HAHammed Adekunle
Hammed Adekunle
UNUrban Nilsson
Urban Nilsson
GÖGöran Örlander
Göran Örlander
EHEmma Holmström
Emma Holmström
We assessed long-term outcomes of a regeneration experiment comparing clearcutting and shelterwood treatments, each with and without mechanical site preparation (MSP). The experiment was established in the early 1990s across Sweden and combined natural regeneration of Scots pine (Pinus sylvestris L.) with planting of Norway spruce (Picea abies (L.) H. Karst.; 2500 seedlings ha–1). The first assessment showed Norway spruce dominance on southern sites and potential for mixed stands in central and northern regions. In 2022–2023, sites were re-measured to assess stand structure and productivity at mid-rotation, and the measured data were then used to parameterize Heureka decision support system (DSS) for simulations to final harvest. Southern sites developed into Norway spruce-dominated stands, with higher volume in clearcut than shelterwood treatments. Central and northern sites developed into mixed species stands, with higher volumes in shelterwood treatments in central Sweden. In northern Sweden, retained shelterwood negatively affected understory growth, highlighting the importance of active management while using shelterwood. Despite these regional differences, land expectation value (LEV) was consistently higher for clearcut treatments. Even in central Sweden, higher volumes in shelterwood treatments did not compensate for the additional costs associated with shelterwood cuttings. Our results show that establishing mixed species stands through shelterwood and planting can be viable depending on site conditions, but likely results in lower revenue than clearcutting.