Earth system models (ESMs) have been developed to represent the role of terrestrial ecosystems on the energy, water, and carbon cycles. However, many ESMs still lack representation of within-ecosystem heterogeneity and diversity. In this paper, we present the Ecosystem Demography model version 2.2 (ED-2.2). In ED-2.2, the biophysical and physiological processes account for the horizontal and vertical heterogeneity of the ecosystem: the energy, water, and carbon cycles are solved separately for a series of vegetation cohorts (groups of individual plants of similar size and plant functional type) distributed across a series of spatially implicit patches (representing collections of micro-environments that have a similar disturbance history). We define the equations that describe the energy, water, and carbon cycles in terms of total energy, water, and carbon, which simplifies the differential equations and guarantees excellent conservation of these quantities in long-term simulation (< 0.1 % error over 50 years). We also show examples of ED-2.2 simulation results at single sites and across tropical South America. These results demonstrate the model's ability to characterize the variability of ecosystem structure, composition, and functioning both at stand and continental scales. A detailed model evaluation was conducted and is presented in a companion paper (Longo et al., 2019a). Finally, we highlight some of the ongoing model developments designed to improve the model's accuracy and performance and to include processes hitherto not represented in the model.

This paper presents a new index to quantify the strength of soil moisture–precipitation coupling in AGCMs and explores how the soil moisture–precipitation coupling in Community Atmosphere Model version 3 (CAM3)–Community Land Model version 3 (CAM3–CLM3) responds to parameterization-induced surface water budget changes. Specifically, this study (a) compares the regions of strong coupling identified by the newly proposed index and the index currently used in the Global Land–Atmosphere Coupling Experiment (GLACE); (b) examines how the surface water budget changes influence the strength of soil moisture–precipitation coupling as measured by the two indexes, respectively; and (c) examines how these changes influence the memory of the coupled land–atmosphere system as measured by the correlation between soil moisture and subsequent precipitation. The new index and the GLACE index are consistent in identifying central North America and West Africa as major regions of strong coupling during June–August (JJA). However, in some areas of western Europe and of subtropical South America where the GLACE index is low, the new index suggests a modest significant coupling during JJA. In response to the surface water budget changes that presumably favor a stronger soil moisture–precipitation coupling, the new index increases, but the GLACE index decreases in a majority of the regions of modest-to-strong coupling, although both show some mixed response. Changes in the land–atmosphere system memory suggest an increase of coupling strength, consistent with results from the new index. The strong dependence of the GLACE index on the relative importance of atmospheric internal variability is identified as a potential cause for the differences between the two indexes. The two indexes emphasize different aspects of soil moisture–precipitation coupling, and one might be more suitable than the other depending on the purpose of individual studies.