Dry-mesic old-growth oak forests are widely distributed remnants across the eastern U.S. and are expected to increase in number and extent as second-growth forests mature. In this study, we synthesize published and unpublished information to better define the species, structure and extent of these forests. Mean site tree density for trees ≥10 cm dbh ranged from 341–620 trees ha−1. In the eastern part of the region, most stand basal areas were >23 m2 ha−1, compared to ≤23 m2 ha−1 in the westernmost stands. Overall, woody species diversity was relatively low compared to old-growth oak forests on moister sites, with tree species per forest ranging from 5–18. The most common species among the stands were white oak (Quercus alba), northern red oak (Quercus rubra), and black oak (Quercus velutina). Shrub and vine species per forest ranged from 1–10, with common species or genera including Virginia creeper (Parthenocissus quinquefolia), poison ivy (Toxicodendron radicans), Vaccinium spp., and grapevines (Vitis spp.). Within the southern Appalachian Mountains, rosebay rhododendron (Rhododendron maximum) and mountain laurel (Kalmia latifolia L.) were common. Herbaceous species per stand ranged from 4–51, with the highest richness occurring in a southern Appalachian oak-hickory forest. The maximum within-stand age of the large trees ranged from 170 to over 365 years. The mean density of standing dead trees ≥10 cm dbh ranged from 31–78 ha−1 and the volume of coarse woody debris ≥10 cm in diameter averaged 52 m3 ha−1. We more fully describe the characteristics of these forests and fill gaps in the collective knowledge of this increasingly important forest type. However, over the past 20 years, there has been scant research on these forests, and older research studies have used a variety of research plots and methods. A uniform approach to surveying these sites is needed to gain a better understanding of these forests before we are faced with caring for an increase in old-growth forest areas.
Nonnative invasive plant species (NNIPS) cause significant damage to the native forest ecosystems in the southern United States forestlands, such as habitat degradation, ecological instability, and biodiversity loss. Taking the state of Alabama as an example, we used more than 5,000 permanent United States Department of Agriculture-Forest Service's Forest Inventory and Analysis (FIA) plots measured between 2001 and 2019 over three measurement cycles to test the suitable modeling unit for quantifying invasion patterns and associated factors for regional NNIPS monitoring and management. NNIPS heavily infest Alabama's forestlands, and forestlands plagued with at least one NNIPS have increased over time: 41.1%, 50.8%, and 54.8% during the past three measurements. Lonicera japonica (Thunb.) was the most abundant NNIPS in Alabama, with at least 26% of its forested lands infested. The FIA data were aggregated with multiple spatial units: five levels of hydrological units, three levels of ecological units, and a county level. Invasion indices were calculated for all spatial units based on NNIPS' presence/absence and average cover in each plot. The best modeling unit was identified based on Moran's test, with the county-level modeling unit providing the best Moran's I value over all measurement periods. Influencing factors of invasion were evaluated based on spatial lag models. Our models show that the invasion index decreased with increases in public forest areas in a county. In contrast, the human population density of neighboring counties positively influenced the invasion index.