Dynamic global vegetation models (DGVMs) suffer insufficiencies in tracking biochemical cycles and ecosystem fluxes. One important reason for these insufficiencies is that DGVMs use fixed parameters (mostly traits) to distinguish attributes and functions of plant functional types (PFTs); however, these traits vary under different climatic conditions. Therefore, it is urgent to quantify trait covariations, including those among specific leaf area (SLA), area-based leaf nitrogen (N_area), and leaf area index (LAI) (in 580 species across 218 sites in this study), and explore new classification methods that can be applied to model vegetation dynamics under future climate change scenarios. We use a redundancy analysis (RDA) to derive trait–climate relationships and employ a Gaussian mixture model (GMM) to project vegetation distributions under different climate scenarios. The results show that (1) the three climatic variables, mean annual temperature (MAT), mean annual precipitation (MAP), and monthly photosynthetically active radiation (mPAR) could capture 65% of the covariations of three functional traits; (2) tropical, subtropical and temperate forest complexes expand while boreal forest, temperate steppe, temperate scrub and tundra shrink under future climate change scenarios; and (3) the GMM classification based on trait covariations should be a powerful candidate for building new generation of DGVM, especially predicting the response of vegetation to future climate changes. This study provides a promising route toward developing reliable, robust and realistic vegetation models and can address a series of limitations in current models.

Shrub layer diversity is an essential component of the forest ecosystem diversity, that contributes significantly to structuring the community and maintaining diversity, especially in plantation forests. In previous studies, researchers have reported the strong relationship among various factors (i.e., soil composition, mean annual temperature, etc.) and shrub diversity. However, how these factors jointly influence shrub diversity and which factors could be considered the key factors is still unknown. In this study, we attempted to quantify the effect among environmental factors, soil factors and forest stand factors on shrub diversity. Twenty-seven variables were selected from 57 Chinese pine plantation plots in Huanglong Mountain, Yanan City, Shaanxi Province, China. The path models showed that latent variable of soil properties is the main effective factor of latent variable of shrub diversity (directly effect, path coefficient = 0.344) and the latent variable of site conditions is another effective factor of latent variable of shrub diversity (indirectly effect, path coefficient = 0.177); Besides, the latent variable of site conditions and forest properties directly affect the latent variable of soil properties (path coefficient = 0.514 and 0.326, respectively). Among the latent variable of soil properties, soil water content (SWC) has the biggest weight of 0.666, which indicated the most significant contribution of SWC to latent variables of shrub diversity. Total nitrogen, weighted 0.375, and total phosphorus, weighted 0.308, are also important factors and make a similar contribution to latent variable of shrub diversity. Soil organic matter (SOM) has a minimal impact (lowest weight, 0.059); among the objective variables of site condition, altitude contributes the most and is followed by litter thickness, weighted at 0.722 and 0.448, respectively. Furthermore, among all the variables affecting the latent variable of forest properties, forest age is recognized as the maximum impactor of soil property change, which weighted −0.941; and is followed by forest stock volume and diameter at breast height (DBH), weighted 0.795 and 0.788, respectively. The crowding index (C) has the lowest weight (−0.235) and demonstrated that spatial distribution and crowding of trees have minimal impact on the latent variable of Soil properties. diversity Overall, our study provides new insights into quantifying the relationships among different driving factors that potentially play a significant role in determining shrub layer diversity within the plantation forest.

Forest carbon density is an important indicator for evaluating forest carbon sink capacities. Accurate carbon density estimation is the basis for studying the response mechanisms of forest ecosystems to global climate change. Airborne light detection and ranging (LiDAR) technology can acquire the vertical structure parameters of forests with a higher precision and penetration ability than traditional optical remote sensing. Combining top of canopy height model (TCH) and allometry models, this paper constructed two prediction models of aboveground carbon density (ACD) with 94 square plots in northwestern China: one model is plot-averaged height-based power model and the other is plot-averaged daisy-chain model. The correlation coefficients (R2) were 0.6725 and 0.6761,which are significantly higher than the correlation coefficients of the traditional percentile model (R2=0.5910).In addition, the correlation between TCH and ACD was significantly better than that between plot-averaged height (AvgH) and ACD, and Lorey’s height (LorH) had no significant correlation with ACD. We also found that plot-level basal area (BA) was a dominant factor in ACD prediction, with a correlation coefficient reaching 0.9182, but this subject requires field investigation. The two models proposed in this study provide a simple and easy approach for estimating ACD in coniferous forests, which can replace the traditional LiDAR percentile method completely.

Competition is an essential driving factor that influences forest community sustainability, yet measuring it poses several challenges. To date, the Competition Index (CI) has generally been the tool of choice for quantifying actual competition. In this study, we proposed using the Total Overlap Index (TOI), a CI in which the Area Overlap (AO) index has been adapted and modified to consider the “shading” and “crowding” effects in the vertical dimension. Next, based on six mixed forest plots in Xiaolong Mountain, Gansu, China, we assessed the results to determine the TOI’s evaluation capability. Individual-tree simulation results showed that compared to the modified Area Overlap index (AO_M), the TOI has superior quantification capability in the vertical direction. The results of the basal area increment (BAI) model showed that the TOI offers the best evaluation capability among the four considered CIs in mixed forest (with Akaike Information Criterion (AIC) of 1041.60 and log-likelihood (LL) of −511.80 in the model fitting test, mean relative error of −28.67%, mean absolute percent error of 117.11%, and root mean square error of 0.7993 in cross-validation). Finally, the TOI was applied in the Kaplan–Meier survival analysis and Cox proportional-hazards analysis. The Kaplan–Meier survival analysis showed a significant difference between the low- (consisting of trees with the TOI lower than 1) and high-competition (consisting of trees with the TOI higher than 1) groups’ survival and hazard curves. Moreover, the results of the Cox proportional-hazards analysis exhibited that the trees in the low-competition group only suffered 34.29% of the hazard risk that trees in the high-competition group suffered. Overall, the TOI expresses more dimensional information than other CIs and appears to be an effective competition index for evaluating individual tree competition. Thus, the competition status quantified using this method may provide new information to guide policy- and decision-makers in sustainable forest management planning projects.

In contrast to intensive management practices focused on wood production, plantations designed to safeguard fragile environments prioritize the sustainable fulfillment of ecological functions. To assess the potential for Chinese pine (Pinus tabuliformis Carr.) plantations in the Loess Hilly Region to effectively serve their ecological protection role over the long term, we selected nine indices representing biological stability, resistance stability, and functional stability. Employing a novel unit circle method, we evaluated the total stability (sum of the three stability components) of 44 plantation plots in Huanglong Mountain. We also explored the connections between total stability and standing spatial structure parameters to offer insights for promptly enhancing stability through thinning. The findings revealed that 79.5% of Chinese pine plantations exhibited moderate total stability, with 20.5% demonstrating good stability. Most plots displayed a random distribution pattern, moderate size differentiation, low species spatial mixing, and high stand crowding. Among the correlations analyzed, mingling exhibited the highest coefficient, followed by differentiation, while the uniform angle index showed the weakest correlation, and crowding displayed an insignificant correlation. While the presence of good functional stability contributed to the moderate total stability, addressing inadequate biological and resistance stability necessitates thinning measures. This study identifies spatial structure types negatively linked to total stability, offering targeted management insights for enhancing the stability of Chinese pine plantations. The stability assessment methodology and indicators presented in this study can serve as a valuable reference for similar plantations with comparable functions and planting conditions.